ORGANOVO HLDGS INC

CUSIP: 68620A104 Class: COM

Invesco Ltd.'s Holding History (CIK: 0000914208)

Note: Option positions (PUT/CALL) are excluded.

Q2 2015
Shares 1,265 Value ($000) $5 Avg Close — Range —
Q1 2014
Shares 14,422 Value ($000) $110 Avg Close — Range —
Q4 2013
Shares 200 Value ($000) $2 Avg Close — Range —