DENALI THERAPEUTICS INC

Ticker: DNLI CUSIP: 24823R105 Class: Common Stock

FIL Ltd's Holding History (CIK: 0000318989)

Note: Option positions (PUT/CALL) are excluded.

Q4 2019

Shares 376,364 Value ($000) $6,556 Avg Close $16.65 Range $14.24 - $20.17

Q3 2019

Shares 376,364 Value ($000) $5,766 Avg Close $19.11 Range $15.29 - $22.26

Q2 2019

Shares 376,364 Value ($000) $7,813 Avg Close $22.41 Range $18.28 - $28.86

Q1 2019

Shares 1,269,036 Value ($000) $29,467 Avg Close $20.92 Range $17.73 - $25.05

Q4 2018

Shares 1,396,885 Value ($000) $28,859 Avg Close $18.14 Range $13.78 - $22.40

Q3 2018

Shares 2,786,979 Value ($000) $60,589 Avg Close $16.54 Range $12.32 - $21.89

Q2 2018

Shares 4,536,715 Value ($000) $69,184 Avg Close $18.84 Range $15.19 - $22.99

Q1 2018

Shares 4,184,982 Value ($000) $82,402 Avg Close $20.56 Range $14.96 - $25.79

Q4 2017

Shares 4,184,982 Value ($000) $65,453 Avg Close $17.97 Range $14.72 - $22.95

Period	Shares	Value ($000)	Average Close	Price Range
Q4 2019	376,364	$6,556	$16.65	$14.24 - $20.17
Q3 2019	376,364	$5,766	$19.11	$15.29 - $22.26
Q2 2019	376,364	$7,813	$22.41	$18.28 - $28.86
Q1 2019	1,269,036	$29,467	$20.92	$17.73 - $25.05
Q4 2018	1,396,885	$28,859	$18.14	$13.78 - $22.40
Q3 2018	2,786,979	$60,589	$16.54	$12.32 - $21.89
Q2 2018	4,536,715	$69,184	$18.84	$15.19 - $22.99
Q1 2018	4,184,982	$82,402	$20.56	$14.96 - $25.79
Q4 2017	4,184,982	$65,453	$17.97	$14.72 - $22.95