DENALI THERAPEUTICS INC

Ticker: DNLI CUSIP: 24823R105 Class: Common Stock

Russell Investments Group, Ltd.'s Holding History (CIK: 0001692234)

Note: Option positions (PUT/CALL) are excluded.

Q4 2025

Shares 13,467 Value ($000) $222 Avg Close $16.88 Range $13.75 - $20.61

Q3 2025

Shares 13,799 Value ($000) $200 Avg Close $14.37 Range $12.58 - $16.20

Q2 2025

Shares 2,558 Value ($000) $36 Avg Close $14.07 Range $10.57 - $16.97

Q1 2025

Shares 2,641 Value ($000) $36 Avg Close $19.16 Range $13.30 - $24.34

Q4 2024

Shares 22,116 Value ($000) $451 Avg Close $25.60 Range $19.93 - $33.33

Q3 2024

Shares 21,974 Value ($000) $640 Avg Close $24.79 Range $19.74 - $32.13

Q2 2024

Shares 22,914 Value ($000) $532 Avg Close $19.20 Range $14.56 - $23.87

Q1 2024

Shares 85,628 Value ($000) $1,757 Avg Close $18.79 Range $15.49 - $23.52

Q4 2023

Shares 111,473 Value ($000) $2,392 Avg Close $19.81 Range $15.45 - $23.81

Q3 2023

Shares 114,058 Value ($000) $2,353 Avg Close $25.37 Range $20.52 - $30.79

Q2 2023

Shares 106,814 Value ($000) $3,152 Avg Close $28.12 Range $22.72 - $33.31

Q1 2023

Shares 82,295 Value ($000) $1,896 Avg Close $27.52 Range $21.74 - $32.81

Q4 2022

Shares 99,373 Value ($000) $2,764 Avg Close $29.75 Range $25.96 - $34.78

Q3 2022

Shares 106,861 Value ($000) $3,277 Avg Close $32.51 Range $25.88 - $39.43

Q2 2022

Shares 88,230 Value ($000) $2,595 Avg Close $26.20 Range $20.24 - $35.62

Q1 2022

Shares 51,520 Value ($000) $1,656 Avg Close $34.41 Range $28.76 - $47.47

Q4 2021

Shares 66,121 Value ($000) $2,952 Avg Close $47.38 Range $41.10 - $56.08

Q3 2021

Shares 65,351 Value ($000) $3,284 Avg Close $56.36 Range $47.02 - $79.50

Q2 2021

Shares 63,757 Value ($000) $4,986 Avg Close $61.86 Range $49.20 - $79.70

Q1 2021

Shares 47,966 Value ($000) $2,726 Avg Close $66.94 Range $52.23 - $84.25

Q4 2020

Shares 80,095 Value ($000) $6,710 Avg Close $60.45 Range $35.89 - $93.94

Q3 2020

Shares 54,599 Value ($000) $1,956 Avg Close $30.12 Range $22.36 - $39.22

Q2 2020

Shares 54,599 Value ($000) $1,319 Avg Close $23.51 Range $15.72 - $29.39

Q1 2020

Shares 71,729 Value ($000) $1,255 Avg Close $20.86 Range $12.39 - $30.41

Q4 2019

Shares 116,919 Value ($000) $2,036 Avg Close $16.65 Range $14.24 - $20.17

Q3 2019

Shares 69,123 Value ($000) $1,060 Avg Close $19.11 Range $15.29 - $22.26

Q2 2019

Shares 35,540 Value ($000) $736 Avg Close $22.41 Range $18.28 - $28.86

Q1 2019

Shares 25,890 Value ($000) $601 Avg Close $20.92 Range $17.73 - $25.05

Q4 2018

Shares 16,796 Value ($000) $347 Avg Close $18.14 Range $13.78 - $22.40

Period	Shares	Value ($000)	Average Close	Price Range
Q4 2025	13,467	$222	$16.88	$13.75 - $20.61
Q3 2025	13,799	$200	$14.37	$12.58 - $16.20
Q2 2025	2,558	$36	$14.07	$10.57 - $16.97
Q1 2025	2,641	$36	$19.16	$13.30 - $24.34
Q4 2024	22,116	$451	$25.60	$19.93 - $33.33
Q3 2024	21,974	$640	$24.79	$19.74 - $32.13
Q2 2024	22,914	$532	$19.20	$14.56 - $23.87
Q1 2024	85,628	$1,757	$18.79	$15.49 - $23.52
Q4 2023	111,473	$2,392	$19.81	$15.45 - $23.81
Q3 2023	114,058	$2,353	$25.37	$20.52 - $30.79
Q2 2023	106,814	$3,152	$28.12	$22.72 - $33.31
Q1 2023	82,295	$1,896	$27.52	$21.74 - $32.81
Q4 2022	99,373	$2,764	$29.75	$25.96 - $34.78
Q3 2022	106,861	$3,277	$32.51	$25.88 - $39.43
Q2 2022	88,230	$2,595	$26.20	$20.24 - $35.62
Q1 2022	51,520	$1,656	$34.41	$28.76 - $47.47
Q4 2021	66,121	$2,952	$47.38	$41.10 - $56.08
Q3 2021	65,351	$3,284	$56.36	$47.02 - $79.50
Q2 2021	63,757	$4,986	$61.86	$49.20 - $79.70
Q1 2021	47,966	$2,726	$66.94	$52.23 - $84.25
Q4 2020	80,095	$6,710	$60.45	$35.89 - $93.94
Q3 2020	54,599	$1,956	$30.12	$22.36 - $39.22
Q2 2020	54,599	$1,319	$23.51	$15.72 - $29.39
Q1 2020	71,729	$1,255	$20.86	$12.39 - $30.41
Q4 2019	116,919	$2,036	$16.65	$14.24 - $20.17
Q3 2019	69,123	$1,060	$19.11	$15.29 - $22.26
Q2 2019	35,540	$736	$22.41	$18.28 - $28.86
Q1 2019	25,890	$601	$20.92	$17.73 - $25.05
Q4 2018	16,796	$347	$18.14	$13.78 - $22.40