IONIS PHARMACEUTICALS INC

Ticker: IONS CUSIP: 462222100 Class: COM

Penserra Capital Management LLC's Holding History (CIK: 0001593324)

Note: Option positions (PUT/CALL) are excluded.

Q4 2016

Shares 471 Value ($000) $22 Avg Close $39.57 Range $24.58 - $57.00

Q3 2016

Shares 390 Value ($000) $14 Avg Close $31.29 Range $23.26 - $40.82

Q2 2016

Shares 234 Value ($000) $5 Avg Close $32.26 Range $19.59 - $46.75

Q1 2016

Shares 150 Value ($000) $6 Avg Close $40.62 Range $30.93 - $62.68

Q4 2015

Shares 118 Value ($000) $7 Avg Close $53.96 Range $38.30 - $65.34

Period	Shares	Value ($000)	Average Close	Price Range
Q4 2016	471	$22	$39.57	$24.58 - $57.00
Q3 2016	390	$14	$31.29	$23.26 - $40.82
Q2 2016	234	$5	$32.26	$19.59 - $46.75
Q1 2016	150	$6	$40.62	$30.93 - $62.68
Q4 2015	118	$7	$53.96	$38.30 - $65.34