IONIS PHARMACEUTICALS INC

Ticker: IONS CUSIP: 462222100 Class: COM

Note: Option positions (PUT/CALL) are excluded.

InstitutionSharesValue ($000)
FMR LLC24,072,101$1,904,344
Capital World Investors17,961,299$1,420,918
VANGUARD GROUP INC16,440,459$1,300,605
BlackRock, Inc.10,647,863$842,352
WELLINGTON MANAGEMENT GROUP LLP6,260,325$495,254
T. Rowe Price Investment Management, Inc.5,789,863$458
Bellevue Group AG4,802,199$379,902
STATE STREET CORP3,974,865$314,452
GEODE CAPITAL MANAGEMENT, LLC3,148,849$249,158
TWO SIGMA ADVISERS, LP3,052,111$241,453
PRICE T ROWE ASSOCIATES INC /MD/2,987,840$236
TWO SIGMA INVESTMENTS, LP2,658,646$210,325
Tweedy, Browne Co LLC2,464,930$195,001
JPMORGAN CHASE & CO2,304,792$182,332
CITADEL ADVISORS LLC2,134,636$168,871
DEERFIELD MANAGEMENT COMPANY, L.P.1,979,669$156,612
BAKER BROS. ADVISORS LP1,791,841$141,753
Deep Track Capital, LP1,700,000$134,487
AMERICAN CENTURY COMPANIES INC1,617,106$127,929
CHARLES SCHWAB INVESTMENT MANAGEMENT INC1,541,841$121,975
SUMMIT PARTNERS PUBLIC ASSET MANAGEMENT, LLC1,462,706$115,715
Pictet Asset Management Holding SA1,418,491$112,476
UBS ASSET MANAGEMENT AMERICAS INC1,388,567$109,850
BNP Paribas Investment Partners S.A.1,352,837$107
Polar Capital Holdings Plc1,333,424$105,487
FRANKLIN RESOURCES INC1,321,345$104,532
Invesco Ltd.1,269,290$100,414
Artisan Partners Limited Partnership1,181,416$93,462
ROYAL BANK OF CANADA1,118,797$88,509
Connor, Clark & Lunn Investment Management Ltd.1,076,370$85,152
ARROWSTREET CAPITAL, LIMITED PARTNERSHIP1,063,327$84,120
UBS Group AG1,045,041$82,673
MORGAN STANLEY1,019,906$80,685
NOMURA ASSET MANAGEMENT INTERNATIONAL INC.864,892$68,422
GOLDMAN SACHS GROUP INC855,966$67,715
NOMURA HOLDINGS INC848,815$67,150
NORTHERN TRUST CORP846,718$66,984
BARCLAYS PLC842,871$66,680
MASSACHUSETTS FINANCIAL SERVICES CO /MA/812,747$64,296
GW&K Investment Management, LLC786,668$62
BANK OF AMERICA CORP /DE/783,579$61,989
TD Asset Management Inc773,184$61,167
GROUPAMA ASSET MANAGMENT766,889$60,669
Candriam S.C.A.677,629$53,610
Nuveen, LLC660,769$52,273
AMERIPRISE FINANCIAL INC598,748$47,367
VOLORIDGE INVESTMENT MANAGEMENT, LLC584,970$46,277
EVENTIDE ASSET MANAGEMENT, LLC575,000$45,488
Tri Locum Partners LP557,174$44,078
Eversept Partners, LP538,218$42,578
Bank of New York Mellon Corp537,886$42,552
FIRST TRUST ADVISORS LP525,239$41,552
Alyeska Investment Group, L.P.481,711$38,108
Hudson Bay Capital Management LP475,000$37,577
PINNACLE ASSOCIATES LTD461,968$36,546
Amundi460,947$36,466
Squarepoint Ops LLC436,279$34,514
ARK Investment Management LLC422,968$33,461
TEACHER RETIREMENT SYSTEM OF TEXAS422,899$33,456
BOONE CAPITAL MANAGEMENT LLC417,833$33,055
C WorldWide Group Holding A/S366,000$29
Jain Global LLC356,259$28,184
ALLIANCEBERNSTEIN L.P.341,849$27,044
BNP PARIBAS ARBITRAGE, SA334,608$26,471
D. E. Shaw & Co., Inc.332,803$26,328
Legal & General Group Plc325,148$25,722
NORGES BANK320,160$25,328
Swiss National Bank298,500$23,614
WELLS FARGO & COMPANY/MN280,045$22,154
Zurich Insurance Group Ltd/FI261,820$20,713
RAYMOND JAMES FINANCIAL INC257,210$20,348
MARSHALL WACE, LLP256,933$20,326
NEW YORK STATE COMMON RETIREMENT FUND241,853$19,133
Privium Fund Management B.V.240,824$19,052
Davern Capital Partners, LP238,477$18,866
Orion Investment Co225,161$17,812
RENAISSANCE TECHNOLOGIES LLC223,607$17,690
Universal- Beteiligungs- und Servicegesellschaft mbH219,574$17,371
DAVENPORT & Co LLC217,897$17,238
Corient Private Wealth LLC214,936$17,004
CALIFORNIA PUBLIC EMPLOYEES RETIREMENT SYSTEM213,368$16,880
TOCQUEVILLE ASSET MANAGEMENT L.P.203,730$16,117
Ikarian Capital, LLC202,038$15,983
STIFEL FINANCIAL CORP198,863$15,732
SECTOR GAMMA AS184,977$14,634
Focus Partners Wealth181,817$14,384
DIMENSIONAL FUND ADVISORS LP176,035$13,925
RHUMBLINE ADVISERS175,387$13,875
DZ BANK AG Deutsche Zentral Genossenschafts Bank, Frankfurt167,657$13,263
Man Group plc166,042$13,136
Sicart Associates LLC163,557$12,939
Rhenman & Partners Asset Management AB158,500$12,539
CenterBook Partners LP153,949$12,179
MANUFACTURERS LIFE INSURANCE COMPANY, THE153,241$12,105
DEUTSCHE BANK AG\149,443$11,822
GREAT LAKES ADVISORS, LLC149,180$11,802
ADAR1 Capital Management, LLC147,130$11,639
Artia Global Partners LP144,376$11,422
CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM144,173$11,406
Fisher Asset Management, LLC141,615$11,203
388 other institutions7,567,750$543,486
Total (488 institutions)166,896,811$12,256,583