PROQR THRAPEUTICS N V

Ticker: PRQR CUSIP: N71542109 Class: SHS EURO

TOCQUEVILLE ASSET MANAGEMENT L.P.'s Holding History (CIK: 0000883961)

Note: Option positions (PUT/CALL) are excluded.

Q1 2016
Shares 21,500 Value ($000) $105 Avg Close $5.08 Range $3.48 - $8.96
Q4 2015
Shares 21,500 Value ($000) $187 Avg Close $11.50 Range $6.95 - $16.23