PROQR THRAPEUTICS N V

Ticker: PRQR CUSIP: N71542109 Class: SHS EURO

Note: Option positions (PUT/CALL) are excluded.

InstitutionSharesValue ($000)
ADAGE CAPITAL PARTNERS GP, L.L.C.6,253,467$12,632
Privium Fund Management B.V.5,357,762$10,823
abrdn plc2,950,676$5,960
Affinity Asset Advisors, LLC2,454,278$4,958
Sio Capital Management, LLC2,164,181$4,372
DAFNA Capital Management LLC1,567,561$3,166
Kynam Capital Management, LP1,566,317$3,164
Ikarian Capital, LLC1,492,229$3,014
MORGAN STANLEY1,367,119$2,762
PLATINUM INVESTMENT MANAGEMENT LTD1,235,514$2,496
ArrowMark Colorado Holdings LLC1,164,722$2,353
M28 Capital Management LP1,019,498$2,059
TWO SIGMA INVESTMENTS, LP839,384$1,696
Point72 Asset Management, L.P.813,650$1,644
UBS Group AG797,664$1,611
NOMURA ASSET MANAGEMENT INTERNATIONAL INC.620,000$1,252
Walleye Capital LLC591,785$1,195
CITADEL ADVISORS LLC583,222$1,178
TWO SIGMA ADVISERS, LP452,400$914
MILLENNIUM MANAGEMENT LLC434,991$879
Monaco Asset Management SAM428,803$866
BOOTHBAY FUND MANAGEMENT, LLC355,283$718
RAYMOND JAMES FINANCIAL INC342,886$693
NORTHERN TRUST CORP294,742$595
Palo Alto Investors LP188,807$381
BANK OF AMERICA CORP /DE/173,924$351
DIADEMA PARTNERS LP167,532$338
JANE STREET GROUP, LLC160,175$324
BNP PARIBAS ARBITRAGE, SA157,149$317
XTX Topco Ltd117,900$238
DEUTSCHE BANK AG\108,762$220
Quadrature Capital Ltd104,231$211
OneDigital Investment Advisors LLC85,000$172
GEODE CAPITAL MANAGEMENT, LLC84,909$172
Lynx1 Capital Management LP68,486$138
Persistent Asset Partners Ltd45,154$91
Velan Capital Investment Management LP45,000$91
SIMPLEX TRADING, LLC44,269$89
GROUP ONE TRADING, L.P.30,388$61
FIFTH THIRD BANCORP30,000$61
RFG Advisory, LLC30,000$61
Invesco Ltd.25,347$51
MOODY LYNN & LIEBERSON, LLC25,000$51
Vanguard Personalized Indexing Management, LLC22,314$45
PNC Financial Services Group, Inc.20,097$41
Keating Financial Advisory Services, Inc.20,000$40
Mariner, LLC19,219$39
GUGGENHEIM CAPITAL LLC17,224$35
Atom Investors LP13,107$26
ROYAL BANK OF CANADA10,303$20
TRUIST FINANCIAL CORP10,000$20
NANO CAP NEW MILLENNIUM GROWTH FUND L P10,000$20
ADAR1 Capital Management, LLC10,000$20
WELLS FARGO & COMPANY/MN4,000$8
BlackRock, Inc.3,945$8
Steward Partners Investment Advisory, LLC1,300$3
Tower Research Capital LLC (TRC)1,211$2
Farther Finance Advisors, LLC1,000$2
PFS Partners, LLC1,000$2
SBI Securities Co., Ltd.515$1
JPMORGAN CHASE & CO283$1
TD Waterhouse Canada Inc.40$0
Rockefeller Capital Management L.P.34$0
Total (63 institutions)37,005,759$74,750