ARCTURUS THERAPEUTICS HLDGS

Ticker: ARCT CUSIP: 03969T109 Class: Common Stock

Note: Option positions (PUT/CALL) are excluded.

InstitutionSharesValue ($000)
FEDERATED HERMES, INC.3,303,640$20,251
BlackRock, Inc.2,619,684$16,059
ARK Investment Management LLC2,611,099$16,006
VANGUARD GROUP INC2,089,641$12,809
Sumitomo Mitsui Trust Holdings, Inc.2,026,588$12,423
Nikko Asset Management Americas, Inc.2,026,588$12,403
GEODE CAPITAL MANAGEMENT, LLC618,134$3,790
STATE STREET CORP564,266$3,459
TWO SIGMA INVESTMENTS, LP546,686$3,351
TWO SIGMA ADVISERS, LP511,043$3,133
DIMENSIONAL FUND ADVISORS LP490,429$3,006
ACADIAN ASSET MANAGEMENT LLC427,894$3
JANE STREET GROUP, LLC422,419$2,589
Empire Life Investments Inc.412,402$2,528
GOLDMAN SACHS GROUP INC391,227$2,398
MILLENNIUM MANAGEMENT LLC304,777$1,868
JACOBS LEVY EQUITY MANAGEMENT, INC293,050$1,796
Hennion & Walsh Asset Management, Inc.240,303$1,473
UBS Group AG227,893$1,397
Point72 Asset Management, L.P.222,163$1,362
MORGAN STANLEY213,392$1,308
JPMORGAN CHASE & CO206,169$1,264
NORTHERN TRUST CORP200,781$1,231
CHARLES SCHWAB INVESTMENT MANAGEMENT INC183,855$1,127
BNP PARIBAS ARBITRAGE, SA146,767$900
RENAISSANCE TECHNOLOGIES LLC119,905$735
Atom Investors LP106,400$652
Invesco Ltd.103,116$632
Susquehanna Portfolio Strategies, LLC103,077$632
D. E. Shaw & Co., Inc.97,823$600
IQ EQ FUND MANAGEMENT (IRELAND) Ltd91,576$561
XTX Topco Ltd82,155$504
Bank of New York Mellon Corp76,301$468
SUSQUEHANNA INTERNATIONAL GROUP, LLP74,729$458
Schonfeld Strategic Advisors LLC69,755$428
Engineers Gate Manager LP68,180$418
Woodline Partners LP66,577$408
AMERICAN CENTURY COMPANIES INC66,485$408
HSBC HOLDINGS PLC66,131$402
MASSACHUSETTS FINANCIAL SERVICES CO /MA/61,357$376
Nuveen, LLC52,222$320
Quantinno Capital Management LP52,123$320
BARCLAYS PLC50,589$310
Kovack Advisors, Inc.46,023$282
Russell Investments Group, Ltd.44,425$272
Jain Global LLC44,085$270
DekaBank Deutsche Girozentrale43,889$1
AQR CAPITAL MANAGEMENT LLC43,403$266
WELLS FARGO & COMPANY/MN41,323$253
PANAGORA ASSET MANAGEMENT INC34,768$213
Squarepoint Ops LLC34,285$210
BANK OF AMERICA CORP /DE/34,088$209
MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.33,172$203
CANADA PENSION PLAN INVESTMENT BOARD30,900$189
RHUMBLINE ADVISERS30,678$188
AXQ CAPITAL, LP28,668$176
Green Alpha Advisors, LLC28,282$173
MGO ONE SEVEN LLC28,117$172
TritonPoint Partners, LLC28,072$172
ALLIANCEBERNSTEIN L.P.27,720$170
Aristides Capital LLC26,283$161
PRICE T ROWE ASSOCIATES INC /MD/26,114$0
Brevan Howard Capital Management LP24,680$151
AlphaQuest LLC23,727$145
ALPS ADVISORS INC23,325$143
nVerses Capital, LLC22,904$140
Freestone Capital Holdings, LLC18,655$114
DEUTSCHE BANK AG\18,373$113
COMMONWEALTH EQUITY SERVICES, LLC18,209$112
Byrne Asset Management LLC18,050$111
Texas Yale Capital Corp.16,000$98
Diversify Advisory Services, LLC15,000$109
TOCQUEVILLE ASSET MANAGEMENT L.P.13,600$83
GREAT WEST LIFE ASSURANCE CO /CAN/12,009$0
INTECH INVESTMENT MANAGEMENT LLC11,382$70
MERCER GLOBAL ADVISORS INC /ADV10,507$64
Formidable Asset Management, LLC10,100$186
MetLife Investment Management, LLC9,357$57
NEW YORK STATE COMMON RETIREMENT FUND8,497$52
PUBLIC EMPLOYEES RETIREMENT SYSTEM OF OHIO7,651$47
Police & Firemen's Retirement System of New Jersey6,957$43
CITIGROUP INC5,731$35
Walleye Trading LLC5,700$35
China Universal Asset Management Co., Ltd.4,630$28
FMR LLC3,236$20
Legal & General Group Plc2,502$15
LOS ANGELES CAPITAL MANAGEMENT LLC2,363$23
Ameritas Investment Partners, Inc.2,317$14
SIGNATUREFD, LLC2,188$13
OSAIC HOLDINGS, INC.1,957$12
ROYAL BANK OF CANADA1,743$10
CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM1,557$10
Mirae Asset Global Investments Co., Ltd.1,381$8
UBS ASSET MANAGEMENT AMERICAS INC1,100$7
Sterling Capital Management LLC961$6
Bank Julius Baer & Co. Ltd, Zurich800$5
AMALGAMATED BANK795$0
PNC Financial Services Group, Inc.619$4
GF FUND MANAGEMENT CO. LTD.577$4
GAMMA Investing LLC462$3
19 other institutions3,002$16
Total (119 institutions)23,696,290$142,256