AUTOLUS THERAPEUTICS PLC

Institution	Shares	Value ($000)
Scientech Research LLC	48,572	$97
TWO SIGMA SECURITIES, LLC	66,579	$132
Arizona PSPRS Trust	150,000	$299
BNP Paribas Investment Partners S.A.	3,361,156	$7
GSA CAPITAL PARTNERS LLP	119,546	$0
Point72 Asset Management, L.P.	305,359	$608
Federation des caisses Desjardins du Quebec	31,328	$62
Icon Wealth Advisors, LLC	10,000	$20
Independent Advisor Alliance	13,885	$28
SageView Advisory Group, LLC	75	$0

Increased Positions (25)

MAK CAPITAL ONE LLC

Value ($000) $59,711 (+40.8%) Shares 30,005,343 (+15.3%)

ARMISTICE CAPITAL, LLC

Value ($000) $34,825 (+37.0%) Shares 17,500,000 (+12.2%)

Caitong International Asset Management Co., Ltd

Value ($000) $245 (+429.8%) Shares 123,279 (+334.0%)

SCHRODER INVESTMENT MANAGEMENT GROUP

Value ($000) $18,694 (+39.4%) Shares 9,489,345 (+12.5%)

Privium Fund Management B.V.

Value ($000) $649 (+29.0%) Shares 326,245 (+5.6%)

UBS Group AG

Value ($000) $1,745 (+160.8%) Shares 877,100 (+113.6%)

Cetera Investment Advisers

Value ($000) $2,189 (+67.6%) Shares 1,099,750 (+37.3%)

SUSQUEHANNA INTERNATIONAL GROUP, LLP

Value ($000) $1,192 (+210.1%) Shares 598,987 (+154.0%)

BlackRock, Inc.

Value ($000) $7,335 (+29.5%) Shares 3,685,825 (+6.1%)

Hennion & Walsh Asset Management, Inc.

Value ($000) $903 (+31.7%) Shares 453,828 (+7.9%)

MORGAN STANLEY

Value ($000) $2,781 (+40.5%) Shares 1,397,251 (+15.1%)

TUDOR INVESTMENT CORP ET AL

Value ($000) $561 (+66.7%) Shares 282,113 (+36.5%)

JPMORGAN CHASE & CO

Value ($000) $1,949 (+32.4%) Shares 979,275 (+8.4%)

SIMPLEX TRADING, LLC

Value ($000) $115 (+523800.0%) Shares 57,919 (+327.4%)

RHUMBLINE ADVISERS

Value ($000) $64 (+140.4%) Shares 32,251 (+96.9%)

Invesco Ltd.

Value ($000) $103 (+65.7%) Shares 51,642 (+35.7%)

MANUFACTURERS LIFE INSURANCE COMPANY, THE

Value ($000) $99 (+58.5%) Shares 49,864 (+29.8%)

HighPoint Advisor Group LLC

Value ($000) $185 (+28.3%) Shares 92,800 (+5.1%)

SIH Partners, LLLP

Value ($000) $30 (+67.4%) Shares 14,887 (+37.1%)

BNP PARIBAS ARBITRAGE, SA

Value ($000) $10 (+118.2%) Shares 4,782 (+78.7%)

Institution	Prev Shares	Curr Shares	Change
MAK CAPITAL ONE LLC	26,017,616	30,005,343	+3,987,727 (+15.3%)
ARMISTICE CAPITAL, LLC	15,600,000	17,500,000	+1,900,000 (+12.2%)
Caitong International Asset Management Co., Ltd	28,406	123,279	+94,873 (+334.0%)
SCHRODER INVESTMENT MANAGEMENT GROUP	8,433,253	9,489,345	+1,056,092 (+12.5%)
Privium Fund Management B.V.	308,837	326,245	+17,408 (+5.6%)
UBS Group AG	410,662	877,100	+466,438 (+113.6%)
Cetera Investment Advisers	801,250	1,099,750	+298,500 (+37.3%)
SUSQUEHANNA INTERNATIONAL GROUP, LLP	235,828	598,987	+363,159 (+154.0%)
BlackRock, Inc.	3,474,775	3,685,825	+211,050 (+6.1%)
Hennion & Walsh Asset Management, Inc.	420,581	453,828	+33,247 (+7.9%)
MORGAN STANLEY	1,213,773	1,397,251	+183,478 (+15.1%)
TUDOR INVESTMENT CORP ET AL	206,653	282,113	+75,460 (+36.5%)
JPMORGAN CHASE & CO	903,068	979,275	+76,207 (+8.4%)
SIMPLEX TRADING, LLC	13,553	57,919	+44,366 (+327.4%)
RHUMBLINE ADVISERS	16,381	32,251	+15,870 (+96.9%)
Invesco Ltd.	38,049	51,642	+13,593 (+35.7%)
MANUFACTURERS LIFE INSURANCE COMPANY, THE	38,419	49,864	+11,445 (+29.8%)
HighPoint Advisor Group LLC	88,300	92,800	+4,500 (+5.1%)
SIH Partners, LLLP	10,860	14,887	+4,027 (+37.1%)
BNP PARIBAS ARBITRAGE, SA	2,676	4,782	+2,106 (+78.7%)

Decreased Positions (28)

Diametric Capital, LP

Value ($000) $398 (-5.1%) Shares 199,922 (-22.3%)

RENAISSANCE TECHNOLOGIES LLC

Value ($000) $6,180 (-22.3%) Shares 3,105,404 (-36.4%)

XTX Topco Ltd

Value ($000) $257 (-38.7%) Shares 129,246 (-49.8%)

Quadrature Capital Ltd

Value ($000) $84 (-85.5%) Shares 42,341 (-88.1%)

CITADEL ADVISORS LLC

Value ($000) $2,308 (-76.6%) Shares 1,159,791 (-80.8%)

Aubrey Capital Management Ltd

Value ($000) $445 (+18.7%) Shares 225,201 (-3.9%)

PRELUDE CAPITAL MANAGEMENT, LLC

Value ($000) $64 (-45.1%) Shares 32,008 (-55.0%)

MARSHALL WACE, LLP

Value ($000) $538 (-48.6%) Shares 270,277 (-57.9%)

GOLDMAN SACHS GROUP INC

Value ($000) $55 (-93.1%) Shares 27,749 (-94.4%)

JANE STREET GROUP, LLC

Value ($000) $1,355 (-4.9%) Shares 681,000 (-22.1%)

LPL Financial LLC

Value ($000) $85 (-61.2%) Shares 42,625 (-68.2%)

BANK OF AMERICA CORP /DE/

Value ($000) $3,902 (+17.9%) Shares 1,960,766 (-3.4%)

WEALTH EFFECTS LLC

Value ($000) $20 (-23.7%) Shares 10,000 (-37.5%)

GEODE CAPITAL MANAGEMENT, LLC

Value ($000) $220 (-17.8%) Shares 110,578 (-32.7%)

R Squared Ltd

Value ($000) $53 (+6.8%) Shares 26,660 (-12.5%)

WELLS FARGO & COMPANY/MN

Value ($000) $392 (+4.1%) Shares 197,083 (-14.7%)

COMMONWEALTH OF PENNSYLVANIA PUBLIC SCHOOL EMPLS RETRMT SYS

Value ($000) $60 (-36.4%) Shares 30,095 (-47.9%)

ROYAL BANK OF CANADA

Value ($000) $20 (-44.4%) Shares 10,000 (-54.3%)

MAI Capital Management

Value ($000) $0 (-94.4%) Shares 189 (-95.4%)

China Universal Asset Management Co., Ltd.

Value ($000) $65 (+13.5%) Shares 32,687 (-7.0%)

Institution	Prev Shares	Curr Shares	Change
Diametric Capital, LP	257,232	199,922	-57,310 (-22.3%)
RENAISSANCE TECHNOLOGIES LLC	4,881,372	3,105,404	-1,775,968 (-36.4%)
XTX Topco Ltd	257,361	129,246	-128,115 (-49.8%)
Quadrature Capital Ltd	357,263	42,341	-314,922 (-88.1%)
CITADEL ADVISORS LLC	6,044,243	1,159,791	-4,884,452 (-80.8%)
Aubrey Capital Management Ltd	234,454	225,201	-9,253 (-3.9%)
PRELUDE CAPITAL MANAGEMENT, LLC	71,161	32,008	-39,153 (-55.0%)
MARSHALL WACE, LLP	641,952	270,277	-371,675 (-57.9%)
GOLDMAN SACHS GROUP INC	493,753	27,749	-466,004 (-94.4%)
JANE STREET GROUP, LLC	874,474	681,000	-193,474 (-22.1%)
LPL Financial LLC	134,025	42,625	-91,400 (-68.2%)
BANK OF AMERICA CORP /DE/	2,029,593	1,960,766	-68,827 (-3.4%)
WEALTH EFFECTS LLC	16,000	10,000	-6,000 (-37.5%)
GEODE CAPITAL MANAGEMENT, LLC	164,314	110,578	-53,736 (-32.7%)
R Squared Ltd	30,476	26,660	-3,816 (-12.5%)
WELLS FARGO & COMPANY/MN	231,132	197,083	-34,049 (-14.7%)
COMMONWEALTH OF PENNSYLVANIA PUBLIC SCHOOL EMPLS RETRMT SYS	57,761	30,095	-27,666 (-47.9%)
ROYAL BANK OF CANADA	21,897	10,000	-11,897 (-54.3%)
MAI Capital Management	4,098	189	-3,909 (-95.4%)
China Universal Asset Management Co., Ltd.	35,157	32,687	-2,470 (-7.0%)

Exited Positions (19)

Affinity Asset Advisors, LLC

Value ($000) $2,853 Shares 1,750,000

Deep Track Capital, LP

Value ($000) $4,187 Shares 2,568,613

Virtus Investment Advisers, LLC

Value ($000) $86 Shares 52,924

Aigen Investment Management, LP

Value ($000) $107 Shares 65,617

TEMA ETFS LLC

Value ($000) $189 Shares 115,823

Perpetual Ltd

Value ($000) $784 Shares 481,081

Graham Capital Management, L.P.

Value ($000) $241 Shares 147,877

Aquatic Capital Management LLC

Value ($000) $171 Shares 104,684

MILLENNIUM MANAGEMENT LLC

Value ($000) $639 Shares 391,833

ALPS ADVISORS INC

Value ($000) $203 Shares 124,606

NEOS Investment Management LLC

Value ($000) $189 Shares 115,823

TWO SIGMA INVESTMENTS, LP

Value ($000) $281 Shares 172,143

PUBLIC EMPLOYEES RETIREMENT SYSTEM OF OHIO

Value ($000) $216 Shares 132,688

Squarepoint Ops LLC

Value ($000) $150 Shares 91,957

Mariner, LLC

Value ($000) $17 Shares 10,322

GROUP ONE TRADING, L.P.

Value ($000) $14 Shares 8,381

BARCLAYS PLC

Value ($000) $4 Shares 993

HRT FINANCIAL LP

Value ($000) $0 Shares 210,172

PRINCIPAL SECURITIES, INC.

Value ($000) $0 Shares 37

Institution	Prev Shares	Prev Value ($000)
Affinity Asset Advisors, LLC	1,750,000	$2,853
Deep Track Capital, LP	2,568,613	$4,187
Virtus Investment Advisers, LLC	52,924	$86
Aigen Investment Management, LP	65,617	$107
TEMA ETFS LLC	115,823	$189
Perpetual Ltd	481,081	$784
Graham Capital Management, L.P.	147,877	$241
Aquatic Capital Management LLC	104,684	$171
MILLENNIUM MANAGEMENT LLC	391,833	$639
ALPS ADVISORS INC	124,606	$203
NEOS Investment Management LLC	115,823	$189
TWO SIGMA INVESTMENTS, LP	172,143	$281
PUBLIC EMPLOYEES RETIREMENT SYSTEM OF OHIO	132,688	$216
Squarepoint Ops LLC	91,957	$150
Mariner, LLC	10,322	$17
GROUP ONE TRADING, L.P.	8,381	$14
BARCLAYS PLC	993	$4
HRT FINANCIAL LP	210,172	$0
PRINCIPAL SECURITIES, INC.	37	$0