QUANTERIX CORP

Institution	Shares	Value ($000)
GSA CAPITAL PARTNERS LLP	121,525	$1
AXQ CAPITAL, LP	11,009	$70
First Eagle Investment Management, LLC	531,427	$3,380
TUDOR INVESTMENT CORP ET AL	169,695	$1,079
Quadrature Capital Ltd	21,707	$138
ExodusPoint Capital Management, LP	24,080	$153
D. E. Shaw & Co., Inc.	45,627	$290
LOS ANGELES CAPITAL MANAGEMENT LLC	34,978	$175
LPL Financial LLC	24,877	$158
RENAISSANCE TECHNOLOGIES LLC	11,800	$75
Farther Finance Advisors, LLC	4,078	$26
Caitong International Asset Management Co., Ltd	763	$5
QUADRANT CAPITAL GROUP LLC	961	$6
Virtu Financial LLC	20,626	$0
UBS ASSET MANAGEMENT AMERICAS INC	471	$3
SIGNATUREFD, LLC	387	$2
NATIONAL BANK OF CANADA /FI/	85	$1
HRT FINANCIAL LP	13,018	$0
ROTHSCHILD INVESTMENT LLC	13	$0

Increased Positions (41)

Blue Water Life Science Advisors, LP

Value ($000) $13,306 (+155.9%) Shares 2,092,147 (+118.4%)

Trium Capital LLP

Value ($000) $5,199 (+39.0%) Shares 817,451 (+18.7%)

Portolan Capital Management, LLC

Value ($000) $28,783 (+24.4%) Shares 4,525,553 (+6.2%)

Empire Financial Management Company, LLC

Value ($000) $3,929 (+59.9%) Shares 617,830 (+36.6%)

Invenomic Capital Management LP

Value ($000) $9,573 (+26.6%) Shares 1,505,208 (+8.1%)

Parkman Healthcare Partners LLC

Value ($000) $2,092 (+52.8%) Shares 328,956 (+30.4%)

WILLIAM BLAIR INVESTMENT MANAGEMENT, LLC

Value ($000) $13,015 (+82.2%) Shares 2,046,449 (+55.6%)

XTX Topco Ltd

Value ($000) $471 (+667.3%) Shares 74,093 (+555.1%)

AMERIPRISE FINANCIAL INC

Value ($000) $31,775 (+37.9%) Shares 4,996,073 (+17.7%)

BlackRock, Inc.

Value ($000) $21,471 (+27.0%) Shares 3,375,910 (+8.4%)

Jump Financial, LLC

Value ($000) $916 (+42.6%) Shares 144,095 (+21.8%)

STATE STREET CORP

Value ($000) $5,682 (+27.7%) Shares 893,401 (+9.0%)

GEODE CAPITAL MANAGEMENT, LLC

Value ($000) $6,832 (+24.3%) Shares 1,073,967 (+6.1%)

Quantinno Capital Management LP

Value ($000) $406 (+171.6%) Shares 63,791 (+131.9%)

FMR LLC

Value ($000) $1,017 (+59.9%) Shares 159,907 (+36.5%)

CHARLES SCHWAB INVESTMENT MANAGEMENT INC

Value ($000) $2,307 (+28.3%) Shares 362,740 (+9.5%)

VANGUARD GROUP INC

Value ($000) $17,031 (+18.5%) Shares 2,677,833 (+1.1%)

BNP PARIBAS ARBITRAGE, SA

Value ($000) $1,071 (+31.2%) Shares 168,430 (+12.0%)

WELLS FARGO & COMPANY/MN

Value ($000) $1,035 (+32.0%) Shares 162,686 (+12.7%)

MORGAN STANLEY

Value ($000) $5,234 (+19.6%) Shares 822,998 (+2.1%)

Institution	Prev Shares	Curr Shares	Change
Blue Water Life Science Advisors, LP	957,747	2,092,147	+1,134,400 (+118.4%)
Trium Capital LLP	688,597	817,451	+128,854 (+18.7%)
Portolan Capital Management, LLC	4,259,851	4,525,553	+265,702 (+6.2%)
Empire Financial Management Company, LLC	452,430	617,830	+165,400 (+36.6%)
Invenomic Capital Management LP	1,393,005	1,505,208	+112,203 (+8.1%)
Parkman Healthcare Partners LLC	252,211	328,956	+76,745 (+30.4%)
WILLIAM BLAIR INVESTMENT MANAGEMENT, LLC	1,315,280	2,046,449	+731,169 (+55.6%)
XTX Topco Ltd	11,310	74,093	+62,783 (+555.1%)
AMERIPRISE FINANCIAL INC	4,243,512	4,996,073	+752,561 (+17.7%)
BlackRock, Inc.	3,112,980	3,375,910	+262,930 (+8.4%)
Jump Financial, LLC	118,352	144,095	+25,743 (+21.8%)
STATE STREET CORP	819,534	893,401	+73,867 (+9.0%)
GEODE CAPITAL MANAGEMENT, LLC	1,011,890	1,073,967	+62,077 (+6.1%)
Quantinno Capital Management LP	27,510	63,791	+36,281 (+131.9%)
FMR LLC	117,133	159,907	+42,774 (+36.5%)
CHARLES SCHWAB INVESTMENT MANAGEMENT INC	331,246	362,740	+31,494 (+9.5%)
VANGUARD GROUP INC	2,647,515	2,677,833	+30,318 (+1.1%)
BNP PARIBAS ARBITRAGE, SA	150,327	168,430	+18,103 (+12.0%)
WELLS FARGO & COMPANY/MN	144,318	162,686	+18,368 (+12.7%)
MORGAN STANLEY	806,263	822,998	+16,735 (+2.1%)

Decreased Positions (43)

PARAGON ASSOCIATES & PARAGON ASSOCIATES II JOINT VENTURE

Value ($000) $159 (-70.9%) Shares 25,000 (-75.2%)

EASTERLY INVESTMENT PARTNERS LLC

Value ($000) $3,922 (-42.3%) Shares 616,681 (-50.8%)

IRONWOOD INVESTMENT MANAGEMENT LLC

Value ($000) $1,096 (+4.0%) Shares 172,330 (-11.2%)

Numerai GP LLC

Value ($000) $189 (-60.4%) Shares 29,762 (-66.2%)

abrdn plc

Value ($000) $662 (-78.4%) Shares 104,039 (-81.6%)

MILLENNIUM MANAGEMENT LLC

Value ($000) $2,561 (-61.0%) Shares 402,631 (-66.7%)

MARSHALL WACE, LLP

Value ($000) $987 (-70.1%) Shares 155,169 (-74.5%)

Mount Lucas Management LP

Value ($000) $359 (+10.3%) Shares 56,500 (-5.8%)

Williams Jones Wealth Management, LLC.

Value ($000) $4,064 (+9.0%) Shares 638,939 (-6.9%)

Susquehanna Portfolio Strategies, LLC

Value ($000) $351 (-33.9%) Shares 55,150 (-43.5%)

DIMENSIONAL FUND ADVISORS LP

Value ($000) $2,477 (-35.5%) Shares 389,466 (-44.9%)

Russell Investments Group, Ltd.

Value ($000) $1,629 (-30.9%) Shares 256,140 (-41.0%)

ENVESTNET ASSET MANAGEMENT INC

Value ($000) $1,604 (-42.7%) Shares 252,211 (-51.1%)

Schonfeld Strategic Advisors LLC

Value ($000) $393 (-47.1%) Shares 61,722 (-54.8%)

ROYAL BANK OF CANADA

Value ($000) $642 (-52.3%) Shares 101,001 (-59.2%)

CITIGROUP INC

Value ($000) $715 (-39.8%) Shares 112,380 (-48.6%)

AQR CAPITAL MANAGEMENT LLC

Value ($000) $417 (-51.4%) Shares 65,522 (-58.5%)

JPMORGAN CHASE & CO

Value ($000) $725 (-41.4%) Shares 114,021 (-49.9%)

TWO SIGMA ADVISERS, LP

Value ($000) $553 (-28.4%) Shares 86,890 (-38.9%)

JANE STREET GROUP, LLC

Value ($000) $441 (-45.7%) Shares 69,383 (-53.7%)

Institution	Prev Shares	Curr Shares	Change
PARAGON ASSOCIATES & PARAGON ASSOCIATES II JOINT VENTURE	100,671	25,000	-75,671 (-75.2%)
EASTERLY INVESTMENT PARTNERS LLC	1,252,184	616,681	-635,503 (-50.8%)
IRONWOOD INVESTMENT MANAGEMENT LLC	194,074	172,330	-21,744 (-11.2%)
Numerai GP LLC	88,064	29,762	-58,302 (-66.2%)
abrdn plc	564,581	104,039	-460,542 (-81.6%)
MILLENNIUM MANAGEMENT LLC	1,209,199	402,631	-806,568 (-66.7%)
MARSHALL WACE, LLP	608,708	155,169	-453,539 (-74.5%)
Mount Lucas Management LP	60,000	56,500	-3,500 (-5.8%)
Williams Jones Wealth Management, LLC.	686,464	638,939	-47,525 (-6.9%)
Susquehanna Portfolio Strategies, LLC	97,669	55,150	-42,519 (-43.5%)
DIMENSIONAL FUND ADVISORS LP	707,071	389,466	-317,605 (-44.9%)
Russell Investments Group, Ltd.	434,060	256,140	-177,920 (-41.0%)
ENVESTNET ASSET MANAGEMENT INC	515,861	252,211	-263,650 (-51.1%)
Schonfeld Strategic Advisors LLC	136,575	61,722	-74,853 (-54.8%)
ROYAL BANK OF CANADA	247,678	101,001	-146,677 (-59.2%)
CITIGROUP INC	218,578	112,380	-106,198 (-48.6%)
AQR CAPITAL MANAGEMENT LLC	157,975	65,522	-92,453 (-58.5%)
JPMORGAN CHASE & CO	227,771	114,021	-113,750 (-49.9%)
TWO SIGMA ADVISERS, LP	142,190	86,890	-55,300 (-38.9%)
JANE STREET GROUP, LLC	149,779	69,383	-80,396 (-53.7%)

Exited Positions (14)

PharVision Advisers, LLC

Value ($000) $196 Shares 36,144

nVerses Capital, LLC

Value ($000) $60 Shares 4,600

R Squared Ltd

Value ($000) $64 Shares 11,771

Atom Investors LP

Value ($000) $106 Shares 19,571

State of Wyoming

Value ($000) $77 Shares 14,109

PRELUDE CAPITAL MANAGEMENT, LLC

Value ($000) $84 Shares 15,393

Quarry LP

Value ($000) $30 Shares 5,548

Quantbot Technologies LP

Value ($000) $47 Shares 8,673

KESTRA PRIVATE WEALTH SERVICES, LLC

Value ($000) $56 Shares 10,294

HSBC HOLDINGS PLC

Value ($000) $56 Shares 10,516

GROUP ONE TRADING, L.P.

Value ($000) $4 Shares 783

NISA INVESTMENT ADVISORS, LLC

Value ($000) $3 Shares 640

Fortitude Family Office, LLC

Value ($000) $1 Shares 270

SIMPLEX TRADING, LLC

Value ($000) $0 Shares 1,563

Institution	Prev Shares	Prev Value ($000)
PharVision Advisers, LLC	36,144	$196
nVerses Capital, LLC	4,600	$60
R Squared Ltd	11,771	$64
Atom Investors LP	19,571	$106
State of Wyoming	14,109	$77
PRELUDE CAPITAL MANAGEMENT, LLC	15,393	$84
Quarry LP	5,548	$30
Quantbot Technologies LP	8,673	$47
KESTRA PRIVATE WEALTH SERVICES, LLC	10,294	$56
HSBC HOLDINGS PLC	10,516	$56
GROUP ONE TRADING, L.P.	783	$4
NISA INVESTMENT ADVISORS, LLC	640	$3
Fortitude Family Office, LLC	270	$1
SIMPLEX TRADING, LLC	1,563	$0