AGENUS INC

Ticker: AGEN CUSIP: 00847G804 Class: Common Stock

Note: Option positions (PUT/CALL) are excluded.

InstitutionSharesValue ($000)
VANGUARD GROUP INC2,830,736$8,889
Siren, L.L.C.1,064,250$3,342
ACADIAN ASSET MANAGEMENT LLC622,484$2
BlackRock, Inc.558,345$1,753
RENAISSANCE TECHNOLOGIES LLC543,546$1,707
CITADEL ADVISORS LLC423,242$1,329
GEODE CAPITAL MANAGEMENT, LLC372,748$1,171
MORGAN STANLEY285,198$896
SUSQUEHANNA INTERNATIONAL GROUP, LLP233,674$734
MILLENNIUM MANAGEMENT LLC225,339$708
Jump Financial, LLC170,407$535
PANAGORA ASSET MANAGEMENT INC163,148$512
MARSHALL WACE, LLP146,800$461
RAYMOND JAMES FINANCIAL INC141,653$445
STATE STREET CORP137,383$431
D. E. Shaw & Co., Inc.127,638$401
Walleye Capital LLC123,458$388
UBS Group AG118,818$373
TWO SIGMA INVESTMENTS, LP117,117$368
683 Capital Management, LLC114,336$359
Man Group plc114,204$359
Point72 Asset Management, L.P.108,220$340
OMERS ADMINISTRATION Corp100,500$316
NORTHERN TRUST CORP89,586$281
JANE STREET GROUP, LLC73,572$231
Connor, Clark & Lunn Investment Management Ltd.67,959$213
CANADA PENSION PLAN INVESTMENT BOARD66,100$208
GOLDMAN SACHS GROUP INC61,749$194
Susquehanna Portfolio Strategies, LLC59,413$187
Squarepoint Ops LLC55,142$173
ExodusPoint Capital Management, LP47,461$149
BARCLAYS PLC46,273$145
Avestar Capital, LLC41,000$129
Qube Research & Technologies Ltd39,895$125
Engineers Gate Manager LP35,279$111
Virtu Financial LLC34,385$0
TWO SIGMA ADVISERS, LP34,245$108
Graham Capital Management, L.P.31,706$100
Ovata Capital Management Ltd30,800$97
CAPTRUST FINANCIAL ADVISORS30,087$94
Key Client Fiduciary Advisors, LLC27,348$86
OSAIC HOLDINGS, INC.27,139$85
HighTower Advisors, LLC25,142$79
PRICE T ROWE ASSOCIATES INC /MD/24,500$0
LPL Financial LLC23,326$73
GROUP ONE TRADING, L.P.22,239$70
Vanguard Personalized Indexing Management, LLC19,831$62
Lighthouse Investment Partners, LLC18,720$59
Quadrature Capital Ltd17,121$54
HSBC HOLDINGS PLC17,056$54
BANK OF MONTREAL /CAN/16,207$51
SHERBROOKE PARK ADVISERS LLC15,390$48
CHARLES SCHWAB INVESTMENT MANAGEMENT INC15,061$47
PRELUDE CAPITAL MANAGEMENT, LLC14,500$46
Redwood Family Wealth LLC14,140$44
BANK OF AMERICA CORP /DE/13,246$42
R Squared Ltd13,129$41
Post Resch Tallon Group Inc.12,750$40
AQR CAPITAL MANAGEMENT LLC11,576$36
Values First Advisors, Inc.11,065$35
NewEdge Advisors, LLC10,596$33
WELLS FARGO & COMPANY/MN10,184$32
MSA Advisors, LLC10,000$31
EP Wealth Advisors, LLC10,000$31
Tower Research Capital LLC (TRC)8,402$26
FMR LLC5,429$17
First Command Advisory Services, Inc.3,530$11
Pinnacle Wealth Planning Services, Inc.3,373$11
ROYAL BANK OF CANADA2,129$7
CoreCap Advisors, LLC2,000$6
CWM, LLC1,917$0
Optiver Holding B.V.1,373$4
CHILTON CAPITAL MANAGEMENT LLC1,058$3
Covestor Ltd1,011$0
NORTHWESTERN MUTUAL WEALTH MANAGEMENT CO633$2
Physician Wealth Advisors, Inc.621$2
Game Plan Financial Advisors, LLC585$2
RHUMBLINE ADVISERS538$2
HUNTINGTON NATIONAL BANK500$2
Russell Investments Group, Ltd.433$1
NISA INVESTMENT ADVISORS, LLC410$1
JPMORGAN CHASE & CO382$1
JONES FINANCIAL COMPANIES LLLP371$1
Rialto Wealth Management, LLC305$1
Fortitude Family Office, LLC296$1
EverSource Wealth Advisors, LLC274$1
Atlantic Union Bankshares Corp250$1
Federation des caisses Desjardins du Quebec100$0
BNP PARIBAS ARBITRAGE, SA96$0
Allworth Financial LP58$0
Root Financial Partners, LLC25$0
IFP Advisors, Inc15$0
HORIZON FINANCIAL SERVICES, LLC9$0
SRS Capital Advisors, Inc.5$0
Parallel Advisors, LLC5$0
SBI Securities Co., Ltd.5$0
Total (96 institutions)10,124,300$29,646