AGENUS INC

Institution	Shares	Value ($000)
683 Capital Management, LLC	114,336	$359
R Squared Ltd	13,129	$41
Ovata Capital Management Ltd	30,800	$97
OMERS ADMINISTRATION Corp	100,500	$316
CITADEL ADVISORS LLC	423,242	$1,329
Man Group plc	114,204	$359
Point72 Asset Management, L.P.	108,220	$340
PRELUDE CAPITAL MANAGEMENT, LLC	14,500	$46
ExodusPoint Capital Management, LP	47,461	$149
Connor, Clark & Lunn Investment Management Ltd.	67,959	$213
CANADA PENSION PLAN INVESTMENT BOARD	66,100	$208
Lighthouse Investment Partners, LLC	18,720	$59
Qube Research & Technologies Ltd	39,895	$125
TWO SIGMA ADVISERS, LP	34,245	$108
Quadrature Capital Ltd	17,121	$54
GROUP ONE TRADING, L.P.	22,239	$70
Pinnacle Wealth Planning Services, Inc.	3,373	$11
Virtu Financial LLC	34,385	$0
PRICE T ROWE ASSOCIATES INC /MD/	24,500	$0
HORIZON FINANCIAL SERVICES, LLC	9	$0

Increased Positions (22)

RENAISSANCE TECHNOLOGIES LLC

Value ($000) $1,707 (+569.6%) Shares 543,546 (+721.1%)

VANGUARD GROUP INC

Value ($000) $8,889 (-10.2%) Shares 2,830,736 (+10.2%)

Jump Financial, LLC

Value ($000) $535 (-1.8%) Shares 170,407 (+20.4%)

PANAGORA ASSET MANAGEMENT INC

Value ($000) $512 (+5.5%) Shares 163,148 (+29.3%)

Squarepoint Ops LLC

Value ($000) $173 (+222.1%) Shares 55,142 (+294.9%)

SUSQUEHANNA INTERNATIONAL GROUP, LLP

Value ($000) $734 (-0.2%) Shares 233,674 (+22.4%)

UBS Group AG

Value ($000) $373 (+18.3%) Shares 118,818 (+45.0%)

MARSHALL WACE, LLP

Value ($000) $461 (+5.7%) Shares 146,800 (+29.6%)

GEODE CAPITAL MANAGEMENT, LLC

Value ($000) $1,171 (-12.4%) Shares 372,748 (+7.4%)

BARCLAYS PLC

Value ($000) $145 (-4.7%) Shares 46,273 (+66.4%)

JANE STREET GROUP, LLC

Value ($000) $231 (+13.7%) Shares 73,572 (+39.4%)

GOLDMAN SACHS GROUP INC

Value ($000) $194 (+10.2%) Shares 61,749 (+35.1%)

TWO SIGMA INVESTMENTS, LP

Value ($000) $368 (-6.7%) Shares 117,117 (+14.4%)

D. E. Shaw & Co., Inc.

Value ($000) $401 (-8.9%) Shares 127,638 (+11.6%)

BlackRock, Inc.

Value ($000) $1,753 (-16.8%) Shares 558,345 (+2.1%)

NORTHERN TRUST CORP

Value ($000) $281 (-10.6%) Shares 89,586 (+9.7%)

LPL Financial LLC

Value ($000) $73 (-1.9%) Shares 23,326 (+20.3%)

Tower Research Capital LLC (TRC)

Value ($000) $26 (+23.6%) Shares 8,402 (+51.5%)

ACADIAN ASSET MANAGEMENT LLC

Value ($000) $2 (+148.5%) Shares 622,484 (+203.5%)

Optiver Holding B.V.

Value ($000) $4 (+5498.7%) Shares 1,373 (+6765.0%)

Institution	Prev Shares	Curr Shares	Change
RENAISSANCE TECHNOLOGIES LLC	66,200	543,546	+477,346 (+721.1%)
VANGUARD GROUP INC	2,569,533	2,830,736	+261,203 (+10.2%)
Jump Financial, LLC	141,494	170,407	+28,913 (+20.4%)
PANAGORA ASSET MANAGEMENT INC	126,184	163,148	+36,964 (+29.3%)
Squarepoint Ops LLC	13,963	55,142	+41,179 (+294.9%)
SUSQUEHANNA INTERNATIONAL GROUP, LLP	190,954	233,674	+42,720 (+22.4%)
UBS Group AG	81,937	118,818	+36,881 (+45.0%)
MARSHALL WACE, LLP	113,313	146,800	+33,487 (+29.6%)
GEODE CAPITAL MANAGEMENT, LLC	346,991	372,748	+25,757 (+7.4%)
BARCLAYS PLC	27,813	46,273	+18,460 (+66.4%)
JANE STREET GROUP, LLC	52,778	73,572	+20,794 (+39.4%)
GOLDMAN SACHS GROUP INC	45,715	61,749	+16,034 (+35.1%)
TWO SIGMA INVESTMENTS, LP	102,353	117,117	+14,764 (+14.4%)
D. E. Shaw & Co., Inc.	114,329	127,638	+13,309 (+11.6%)
BlackRock, Inc.	547,112	558,345	+11,233 (+2.1%)
NORTHERN TRUST CORP	81,698	89,586	+7,888 (+9.7%)
LPL Financial LLC	19,383	23,326	+3,943 (+20.3%)
Tower Research Capital LLC (TRC)	5,545	8,402	+2,857 (+51.5%)
ACADIAN ASSET MANAGEMENT LLC	205,091	622,484	+417,393 (+203.5%)
Optiver Holding B.V.	20	1,373	+1,353 (+6765.0%)

Decreased Positions (28)

Post Resch Tallon Group Inc.

Value ($000) $40 (-48.0%) Shares 12,750 (-36.3%)

Key Client Fiduciary Advisors, LLC

Value ($000) $86 (-43.1%) Shares 27,348 (-30.3%)

Susquehanna Portfolio Strategies, LLC

Value ($000) $187 (-71.2%) Shares 59,413 (-64.7%)

Walleye Capital LLC

Value ($000) $388 (-67.6%) Shares 123,458 (-60.3%)

BANK OF AMERICA CORP /DE/

Value ($000) $42 (-97.1%) Shares 13,246 (-96.4%)

MORGAN STANLEY

Value ($000) $896 (-57.6%) Shares 285,198 (-48.1%)

MILLENNIUM MANAGEMENT LLC

Value ($000) $708 (-47.3%) Shares 225,339 (-35.4%)

Graham Capital Management, L.P.

Value ($000) $100 (-52.8%) Shares 31,706 (-42.2%)

SHERBROOKE PARK ADVISERS LLC

Value ($000) $48 (-24.2%) Shares 15,390 (-7.0%)

NewEdge Advisors, LLC

Value ($000) $33 (-80.6%) Shares 10,596 (-76.2%)

Values First Advisors, Inc.

Value ($000) $35 (-32.3%) Shares 11,065 (-17.0%)

JPMORGAN CHASE & CO

Value ($000) $1 (-98.9%) Shares 382 (-98.7%)

AQR CAPITAL MANAGEMENT LLC

Value ($000) $36 (-65.4%) Shares 11,576 (-57.6%)

STATE STREET CORP

Value ($000) $431 (-25.6%) Shares 137,383 (-8.8%)

ROYAL BANK OF CANADA

Value ($000) $7 (-85.7%) Shares 2,129 (-83.4%)

OSAIC HOLDINGS, INC.

Value ($000) $85 (-39.4%) Shares 27,139 (-20.1%)

RAYMOND JAMES FINANCIAL INC

Value ($000) $445 (-20.7%) Shares 141,653 (-2.8%)

WELLS FARGO & COMPANY/MN

Value ($000) $32 (-37.7%) Shares 10,184 (-23.5%)

Russell Investments Group, Ltd.

Value ($000) $1 (-89.0%) Shares 433 (-86.5%)

SRS Capital Advisors, Inc.

Value ($000) $0 (-99.4%) Shares 5 (-99.3%)

Institution	Prev Shares	Curr Shares	Change
Post Resch Tallon Group Inc.	20,000	12,750	-7,250 (-36.3%)
Key Client Fiduciary Advisors, LLC	39,233	27,348	-11,885 (-30.3%)
Susquehanna Portfolio Strategies, LLC	168,221	59,413	-108,808 (-64.7%)
Walleye Capital LLC	310,788	123,458	-187,330 (-60.3%)
BANK OF AMERICA CORP /DE/	369,314	13,246	-356,068 (-96.4%)
MORGAN STANLEY	549,126	285,198	-263,928 (-48.1%)
MILLENNIUM MANAGEMENT LLC	348,754	225,339	-123,415 (-35.4%)
Graham Capital Management, L.P.	54,820	31,706	-23,114 (-42.2%)
SHERBROOKE PARK ADVISERS LLC	16,557	15,390	-1,167 (-7.0%)
NewEdge Advisors, LLC	44,446	10,596	-33,850 (-76.2%)
Values First Advisors, Inc.	13,335	11,065	-2,270 (-17.0%)
JPMORGAN CHASE & CO	29,080	382	-28,698 (-98.7%)
AQR CAPITAL MANAGEMENT LLC	27,279	11,576	-15,703 (-57.6%)
STATE STREET CORP	150,583	137,383	-13,200 (-8.8%)
ROYAL BANK OF CANADA	12,837	2,129	-10,708 (-83.4%)
OSAIC HOLDINGS, INC.	33,965	27,139	-6,826 (-20.1%)
RAYMOND JAMES FINANCIAL INC	145,760	141,653	-4,107 (-2.8%)
WELLS FARGO & COMPANY/MN	13,321	10,184	-3,137 (-23.5%)
Russell Investments Group, Ltd.	3,217	433	-2,784 (-86.5%)
SRS Capital Advisors, Inc.	667	5	-662 (-99.3%)

Exited Positions (10)

Torrey Growth & Income Advisors

Value ($000) $507 Shares 131,677

Traynor Capital Management, Inc.

Value ($000) $50 Shares 13,000

nVerses Capital, LLC

Value ($000) $7 Shares 1,300

ARROWSTREET CAPITAL, LIMITED PARTNERSHIP

Value ($000) $120 Shares 31,054

COMERICA BANK

Value ($000) $4 Shares 1,058

CITIGROUP INC

Value ($000) $2 Shares 547

PNC Financial Services Group, Inc.

Value ($000) $0 Shares 63

SIMPLEX TRADING, LLC

Value ($000) $0 Shares 23,075

Advisory Services Network, LLC

Value ($000) $0 Shares 5

Front Row Advisors LLC

Value ($000) $0 Shares 1

Institution	Prev Shares	Prev Value ($000)
Torrey Growth & Income Advisors	131,677	$507
Traynor Capital Management, Inc.	13,000	$50
nVerses Capital, LLC	1,300	$7
ARROWSTREET CAPITAL, LIMITED PARTNERSHIP	31,054	$120
COMERICA BANK	1,058	$4
CITIGROUP INC	547	$2
PNC Financial Services Group, Inc.	63	$0
SIMPLEX TRADING, LLC	23,075	$0
Advisory Services Network, LLC	5	$0
Front Row Advisors LLC	1	$0