| BlackRock, Inc. | 5,395,232 | $656,330 |
| FMR LLC | 5,077,036 | $617,621 |
| VANGUARD GROUP INC | 3,605,279 | $438,582 |
| STATE STREET CORP | 1,275,133 | $155,120 |
| UBS ASSET MANAGEMENT AMERICAS INC | 996,878 | $121,270 |
| GOLDMAN SACHS GROUP INC | 965,931 | $117,506 |
| NOMURA ASSET MANAGEMENT INTERNATIONAL INC. | 832,241 | $101,242 |
| GEODE CAPITAL MANAGEMENT, LLC | 816,474 | $99,339 |
| DRIEHAUS CAPITAL MANAGEMENT LLC | 712,041 | $86,620 |
| Hood River Capital Management LLC | 690,357 | $83,982 |
| Invesco Ltd. | 684,574 | $83,278 |
| MORGAN STANLEY | 667,761 | $81,233 |
| UBS Group AG | 659,296 | $80,203 |
| PRICE T ROWE ASSOCIATES INC /MD/ | 569,970 | $69 |
| EVENTIDE ASSET MANAGEMENT, LLC | 548,267 | $66,698 |
| DIMENSIONAL FUND ADVISORS LP | 489,710 | $59,574 |
| TWO SIGMA INVESTMENTS, LP | 435,969 | $53,036 |
| WELLINGTON MANAGEMENT GROUP LLP | 433,321 | $52,713 |
| JPMORGAN CHASE & CO | 375,261 | $45,651 |
| TWO SIGMA ADVISERS, LP | 362,300 | $44,074 |
| EMERALD ADVISERS, LLC | 351,349 | $42,742 |
| NORTHERN TRUST CORP | 339,216 | $41,266 |
| WASATCH ADVISORS LP | 331,848 | $40,369 |
| CHARLES SCHWAB INVESTMENT MANAGEMENT INC | 323,026 | $39,296 |
| BAILLIE GIFFORD & CO | 316,473 | $38,499 |
| BRAIDWELL LP | 315,341 | $38,361 |
| FRONTIER CAPITAL MANAGEMENT CO LLC | 302,760 | $36,831 |
| BANK OF AMERICA CORP /DE/ | 279,303 | $33,977 |
| WELLS FARGO & COMPANY/MN | 258,550 | $31,453 |
| ROYCE & ASSOCIATES LP | 243,149 | $29,579 |
| EMERALD MUTUAL FUND ADVISERS TRUST | 229,901 | $27,967 |
| Capital World Investors | 227,441 | $27,668 |
| Bank of New York Mellon Corp | 218,596 | $26,592 |
| LORD, ABBETT & CO. LLC | 216,873 | $26 |
| BNP PARIBAS ARBITRAGE, SA | 209,529 | $25,489 |
| BARCLAYS PLC | 199,705 | $24,294 |
| PRINCIPAL FINANCIAL GROUP INC | 193,777 | $23,573 |
| MARSHALL WACE, LLP | 190,470 | $23,171 |
| ArrowMark Colorado Holdings LLC | 186,979 | $22,746 |
| DEUTSCHE BANK AG\ | 164,573 | $20,020 |
| MILLENNIUM MANAGEMENT LLC | 149,031 | $18,130 |
| DAVIDSON KEMPNER CAPITAL MANAGEMENT LP | 146,400 | $17,810 |
| BROWN CAPITAL MANAGEMENT LLC | 145,918 | $17,751 |
| CITADEL ADVISORS LLC | 143,247 | $17,426 |
| BLAIR WILLIAM & CO/IL | 135,889 | $16,531 |
| AMERIPRISE FINANCIAL INC | 124,339 | $15,126 |
| BANK OF MONTREAL /CAN/ | 124,186 | $15,107 |
| Public Sector Pension Investment Board | 118,256 | $14,386 |
| OPPENHEIMER & CO INC | 116,566 | $14,180 |
| RAYMOND JAMES FINANCIAL INC | 112,532 | $13,706 |
| Squarepoint Ops LLC | 110,496 | $13,442 |
| Candriam S.C.A. | 109,186 | $13,283 |
| STATE OF MICHIGAN RETIREMENT SYSTEM | 108,000 | $13,138 |
| Lagoda Investment Management, L.P. | 104,800 | $12,749 |
| PANAGORA ASSET MANAGEMENT INC | 102,025 | $12,411 |
| Montanaro Asset Management Ltd | 101,939 | $12,401 |
| NEXT CENTURY GROWTH INVESTORS LLC | 98,468 | $11,979 |
| Bellevue Group AG | 97,885 | $11,908 |
| Portolan Capital Management, LLC | 96,893 | $11,787 |
| Man Group plc | 95,664 | $11,638 |
| Trexquant Investment LP | 92,481 | $11,250 |
| Universal- Beteiligungs- und Servicegesellschaft mbH | 92,446 | $11,246 |
| Schonfeld Strategic Advisors LLC | 91,065 | $11,078 |
| RHUMBLINE ADVISERS | 88,036 | $10,710 |
| CREDIT AGRICOLE S A | 87,219 | $10,610 |
| Legal & General Group Plc | 85,868 | $10,446 |
| Ensign Peak Advisors, Inc | 80,603 | $9,805 |
| RENAISSANCE GROUP LLC | 77,229 | $9,395 |
| Nuveen, LLC | 75,163 | $9,144 |
| FirstWave Capital Management LLC | 74,562 | $9,070 |
| CITIGROUP INC | 72,044 | $8,764 |
| JANUS HENDERSON GROUP PLC | 71,951 | $8,754 |
| F/M Investments LLC | 65,539 | $7,973 |
| Swiss National Bank | 63,900 | $7,773 |
| ROYAL BANK OF CANADA | 62,833 | $7,643 |
| PEREGRINE CAPITAL MANAGEMENT LLC | 60,157 | $7,318 |
| Bastion Asset Management Inc. | 57,596 | $7,229 |
| Biondo Investment Advisors, LLC | 57,215 | $6,960 |
| PRUDENTIAL FINANCIAL INC | 57,105 | $6,947 |
| Lisanti Capital Growth, LLC | 56,751 | $6,904 |
| SEI INVESTMENTS CO | 55,291 | $6,726 |
| LAZARD ASSET MANAGEMENT LLC | 51,409 | $6,254 |
| FEDERATED HERMES, INC. | 47,301 | $5,754 |
| Parkman Healthcare Partners LLC | 45,174 | $5,495 |
| CANADA PENSION PLAN INVESTMENT BOARD | 44,600 | $5,426 |
| ENVESTNET ASSET MANAGEMENT INC | 43,187 | $5,254 |
| ALLIANCEBERNSTEIN L.P. | 42,142 | $5,127 |
| Gotham Asset Management, LLC | 40,872 | $4,972 |
| Meridiem Capital Partners LP | 36,668 | $4,461 |
| BESSEMER GROUP INC | 33,009 | $4 |
| Mitsubishi UFJ Kokusai Asset Management Co., Ltd. | 32,909 | $4,038 |
| NORDEA INVESTMENT MANAGEMENT AB | 31,992 | $3,953 |
| STATE OF WISCONSIN INVESTMENT BOARD | 31,950 | $3,887 |
| LPL Financial LLC | 31,263 | $3,803 |
| CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM | 30,821 | $3,749 |
| NORGES BANK | 30,574 | $3,719 |
| ALGERT GLOBAL LLC | 30,294 | $4 |
| FRED ALGER MANAGEMENT, LLC | 30,269 | $3,682 |
| MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd. | 29,841 | $3,630 |
| NEW YORK STATE TEACHERS RETIREMENT SYSTEM | 29,503 | $4 |
| 323 other institutions | 1,636,467 | $172,967 |