ATYR PHARMA INC

Ticker: ATYR CUSIP: 002120202 Class: Common Stock

Note: Option positions (PUT/CALL) are excluded.

InstitutionSharesValue ($000)
FMR LLC14,697,995$11,510
FEDERATED HERMES, INC.10,509,678$8,230
BlackRock, Inc.6,549,699$5,129
VANGUARD GROUP INC4,961,871$3,885
GSA CAPITAL PARTNERS LLP2,630,638$2
GEODE CAPITAL MANAGEMENT, LLC2,283,717$1,789
STATE STREET CORP1,724,843$1,351
UBS Group AG1,625,297$1,273
MORGAN STANLEY1,403,009$1,099
Rosalind Advisors, Inc.1,039,080$13,602
TWO SIGMA ADVISERS, LP841,350$659
AQR CAPITAL MANAGEMENT LLC769,820$603
SUSQUEHANNA INTERNATIONAL GROUP, LLP764,743$599
NORTHERN TRUST CORP702,186$550
Tikvah Management LLC460,833$361
TWO SIGMA INVESTMENTS, LP454,443$356
GOLDMAN SACHS GROUP INC345,587$271
Nuveen, LLC227,941$179
CHARLES SCHWAB INVESTMENT MANAGEMENT INC221,767$174
Bank of New York Mellon Corp193,532$152
GROUP ONE TRADING, L.P.191,048$150
JANE STREET GROUP, LLC188,149$147
VICTORY CAPITAL MANAGEMENT INC181,966$142
BARCLAYS PLC174,182$136
BANK OF AMERICA CORP /DE/126,700$99
DEUTSCHE BANK AG\120,341$94
WELLS FARGO & COMPANY/MN116,717$91
JPMORGAN CHASE & CO110,373$86
RHUMBLINE ADVISERS99,762$78
DIMENSIONAL FUND ADVISORS LP97,497$76
XTX Topco Ltd79,365$62
Virtu Financial LLC78,953$0
ALLIANCEBERNSTEIN L.P.64,480$50
KINGSWOOD WEALTH ADVISORS, LLC52,000$41
ADAR1 Capital Management, LLC50,685$40
Invesco Ltd.46,233$36
PRICE T ROWE ASSOCIATES INC /MD/46,126$0
MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.41,844$33
TWO SIGMA SECURITIES, LLC39,006$31
SEI INVESTMENTS CO33,829$26
INTECH INVESTMENT MANAGEMENT LLC31,774$25
LPL Financial LLC30,100$24
Rockefeller Capital Management L.P.28,118$22
NEW YORK STATE COMMON RETIREMENT FUND27,500$22
BNP PARIBAS ARBITRAGE, SA26,518$21
SIMPLEX TRADING, LLC26,227$21
Police & Firemen's Retirement System of New Jersey26,081$20
SG Americas Securities, LLC24,032$0
NEUBERGER BERMAN GROUP LLC20,454$16
RBF Capital, LLC20,000$16
ProShare Advisors LLC17,524$14
Voya Investment Management LLC16,594$13
HSBC HOLDINGS PLC16,136$12
RAYMOND JAMES FINANCIAL INC15,961$12
MAI Capital Management15,239$12
Tower Research Capital LLC (TRC)13,729$11
Apollon Wealth Management, LLC12,900$10
Crossmark Global Holdings, Inc.12,666$10
Gladstone Institutional Advisory LLC11,300$9
R Squared Ltd11,255$9
Belpointe Asset Management LLC10,438$8
Legal & General Group Plc9,570$7
Ameritas Investment Partners, Inc.8,881$7
CWM, LLC7,978$0
Russell Investments Group, Ltd.7,205$6
CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM5,901$5
ROYAL BANK OF CANADA5,253$5
FNY Investment Advisers, LLC4,800$0
Newbridge Financial Services Group, Inc.3,600$3
CITIGROUP INC3,070$2
AMALGAMATED BANK3,014$0
Farther Finance Advisors, LLC2,225$2
JONES FINANCIAL COMPANIES LLLP1,500$1
Sterling Capital Management LLC934$1
PNC Financial Services Group, Inc.478$0
MCF Advisors LLC400$0
NATIXIS280$0
EverSource Wealth Advisors, LLC161$0
Larson Financial Group LLC150$0
ROTHSCHILD INVESTMENT LLC109$0
Caitong International Asset Management Co., Ltd67$0
CALDWELL SUTTER CAPITAL, INC.52$0
Total (82 institutions)54,827,459$53,536