HARVARD BIOSCIENCE INC

Ticker: HBIO CUSIP: 416906105 Class: COM

Note: Option positions (PUT/CALL) are excluded.

InstitutionSharesValue ($000)
WEBER CAPITAL MANAGEMENT LLC /ADV3,200,380$2,140
VANGUARD GROUP INC1,923,275$1,287
ACADIAN ASSET MANAGEMENT LLC1,772,627$1
Meros Investment Management, LP1,217,628$814
CORSAIR CAPITAL MANAGEMENT, L.P.1,108,692$741
RENAISSANCE TECHNOLOGIES LLC1,021,241$683
TWO SIGMA INVESTMENTS, LP1,012,417$677
TWO SIGMA ADVISERS, LP878,556$588
BlackRock, Inc.622,296$416
NANO CAP NEW MILLENNIUM GROWTH FUND L P600,000$401
MILLENNIUM MANAGEMENT LLC546,067$365
GEODE CAPITAL MANAGEMENT, LLC460,793$308
JANE STREET GROUP, LLC415,159$278
Portolan Capital Management, LLC409,650$274
Williams & Novak, LLC348,800$233
CITIGROUP INC193,828$130
ESSEX INVESTMENT MANAGEMENT CO LLC184,678$124
RBF Capital, LLC160,000$107
STATE STREET CORP149,352$100
SUSQUEHANNA INTERNATIONAL GROUP, LLP139,869$94
Assenagon Asset Management S.A.139,218$93
Bank of New York Mellon Corp126,223$84
NORTHERN TRUST CORP91,973$62
Virtu Financial LLC59,508$0
GAMCO INVESTORS, INC. ET AL57,000$38
GOLDMAN SACHS GROUP INC49,751$33
Pasadena Private Wealth, LLC45,000$30
Legato Capital Management LLC43,467$29
CITADEL ADVISORS LLC42,992$29
XTX Topco Ltd39,349$26
GABELLI FUNDS LLC39,000$26
FMR LLC32,899$22
Callan Family Office, LLC29,791$20
DIMENSIONAL FUND ADVISORS LP27,792$19
GREENWOOD CAPITAL ASSOCIATES LLC27,754$19
GROUP ONE TRADING, L.P.25,513$17
Abel Hall, LLC25,283$17
UBS Group AG21,213$14
TWO SIGMA SECURITIES, LLC17,549$12
Qube Research & Technologies Ltd17,029$11
MORGAN STANLEY16,686$11
Vanguard Personalized Indexing Management, LLC16,542$11
Point72 Asset Management, L.P.14,717$10
SIMPLEX TRADING, LLC13,790$9
CERTUITY, LLC13,677$9
GSK plc11,151$7
Ethic Inc.10,942$7
Balboa Wealth Partners10,000$7
SBI Securities Co., Ltd.8,670$6
ROYAL BANK OF CANADA4,795$3
Brown Shipley& Co Ltd4,783$3
Tower Research Capital LLC (TRC)2,395$2
NISA INVESTMENT ADVISORS, LLC2,289$2
JPMORGAN CHASE & CO1,960$1
FEDERATED HERMES, INC.1,656$1
BNP PARIBAS ARBITRAGE, SA680$0
BANK OF AMERICA CORP /DE/451$0
Financial Connections Group, Inc.300$0
PNC Financial Services Group, Inc.144$0
SHP Wealth Management100$0
CWM, LLC54$0
BARCLAYS PLC21$0
Optiver Holding B.V.14$0
Total (63 institutions)17,459,429$10,453