WESBANCO INC

Institution	Shares	Value ($000)
Rhino Investment Partners, Inc	593,025	$19,712
Aristotle Capital Boston, LLC	339,308	$11,279
Watershed Asset Management, L.L.C.	12,739	$423
Gator Capital Management, LLC	43,970	$1,462
Fieldview Capital Management, LLC	16,052	$534
Abel Hall, LLC	6,034	$201
CenterBook Partners LP	26,289	$874
Clear Harbor Asset Management, LLC	15,280	$508
PARTHENON LLC	7,960	$265
RITHOLTZ WEALTH MANAGEMENT	21,696	$721
BEESE FULMER INVESTMENT MANAGEMENT, INC.	6,457	$215
Weiss Asset Management LP	12,444	$414
DRW Securities, LLC	8,081	$269
Vise Technologies, Inc.	6,718	$223
Jain Global LLC	19,688	$654
GSA CAPITAL PARTNERS LLP	8,548	$0
HSBC HOLDINGS PLC	44,800	$1,492
Brevan Howard Capital Management LP	9,781	$325
Waverly Advisors, LLC	6,647	$221
Merit Financial Group, LLC	6,593	$219

Increased Positions (75)

WESBANCO BANK INC

Value ($000) $52,196 (+9.8%) Shares 1,570,275 (+5.5%)

1492 Capital Management LLC

Value ($000) $2,355 (+7.4%) Shares 70,850 (+3.2%)

Legato Capital Management LLC

Value ($000) $1,504 (+87.1%) Shares 45,238 (+79.7%)

Edgestream Partners, L.P.

Value ($000) $4,852 (+36.2%) Shares 145,966 (+30.8%)

Y-Intercept (Hong Kong) Ltd

Value ($000) $2,478 (+150.5%) Shares 74,555 (+140.6%)

BRIDGEWAY CAPITAL MANAGEMENT, LLC

Value ($000) $18,411 (+9.4%) Shares 553,881 (+5.1%)

TWO SIGMA INVESTMENTS, LP

Value ($000) $15,596 (+120.9%) Shares 469,183 (+112.2%)

SCHULHOFF & CO INC

Value ($000) $1,405 (+6.0%) Shares 42,275 (+1.8%)

Hood River Capital Management LLC

Value ($000) $29,731 (+6.9%) Shares 894,431 (+2.7%)

Atom Investors LP

Value ($000) $2,474 (+13.4%) Shares 74,441 (+8.9%)

TWO SIGMA ADVISERS, LP

Value ($000) $5,387 (+172.2%) Shares 162,060 (+161.5%)

Verition Fund Management LLC

Value ($000) $62,230 (+5.3%) Shares 1,872,132 (+1.2%)

WELLINGTON MANAGEMENT GROUP LLP

Value ($000) $91,883 (+18.3%) Shares 2,764,222 (+13.7%)

GOLDMAN SACHS GROUP INC

Value ($000) $53,441 (+35.8%) Shares 1,607,740 (+30.5%)

Russell Investments Group, Ltd.

Value ($000) $8,696 (+55.3%) Shares 261,613 (+49.2%)

RAYMOND JAMES FINANCIAL INC

Value ($000) $18,421 (+46.8%) Shares 554,186 (+41.0%)

Boston Partners

Value ($000) $5,338 (+84.1%) Shares 160,593 (+76.9%)

Qube Research & Technologies Ltd

Value ($000) $12,565 (+22.9%) Shares 378,000 (+18.1%)

VANGUARD GROUP INC

Value ($000) $326,399 (+6.5%) Shares 9,819,476 (+2.3%)

SEI INVESTMENTS CO

Value ($000) $16,225 (+17.3%) Shares 488,113 (+12.7%)

Institution	Prev Shares	Curr Shares	Change
WESBANCO BANK INC	1,488,301	1,570,275	+81,974 (+5.5%)
1492 Capital Management LLC	68,681	70,850	+2,169 (+3.2%)
Legato Capital Management LLC	25,171	45,238	+20,067 (+79.7%)
Edgestream Partners, L.P.	111,595	145,966	+34,371 (+30.8%)
Y-Intercept (Hong Kong) Ltd	30,988	74,555	+43,567 (+140.6%)
BRIDGEWAY CAPITAL MANAGEMENT, LLC	527,148	553,881	+26,733 (+5.1%)
TWO SIGMA INVESTMENTS, LP	221,151	469,183	+248,032 (+112.2%)
SCHULHOFF & CO INC	41,535	42,275	+740 (+1.8%)
Hood River Capital Management LLC	871,211	894,431	+23,220 (+2.7%)
Atom Investors LP	68,331	74,441	+6,110 (+8.9%)
TWO SIGMA ADVISERS, LP	61,980	162,060	+100,080 (+161.5%)
Verition Fund Management LLC	1,850,511	1,872,132	+21,621 (+1.2%)
WELLINGTON MANAGEMENT GROUP LLP	2,431,637	2,764,222	+332,585 (+13.7%)
GOLDMAN SACHS GROUP INC	1,232,456	1,607,740	+375,284 (+30.5%)
Russell Investments Group, Ltd.	175,381	261,613	+86,232 (+49.2%)
RAYMOND JAMES FINANCIAL INC	392,978	554,186	+161,208 (+41.0%)
Boston Partners	90,787	160,593	+69,806 (+76.9%)
Qube Research & Technologies Ltd	320,088	378,000	+57,912 (+18.1%)
VANGUARD GROUP INC	9,595,369	9,819,476	+224,107 (+2.3%)
SEI INVESTMENTS CO	433,025	488,113	+55,088 (+12.7%)

Decreased Positions (92)

FJ Capital Management LLC

Value ($000) $11,539 (-44.8%) Shares 347,136 (-46.9%)

MALTESE CAPITAL MANAGEMENT LLC

Value ($000) $9,141 (-28.8%) Shares 275,000 (-31.6%)

QSV Equity Investors LLC

Value ($000) $1,138 (-30.8%) Shares 34,246 (-33.5%)

BRANT POINT INVESTMENT MANAGEMENT LLC

Value ($000) $6,784 (-27.8%) Shares 204,105 (-30.6%)

Bridge City Capital, LLC

Value ($000) $2,749 (-10.3%) Shares 82,700 (-13.8%)

BFSG, LLC

Value ($000) $322 (-77.5%) Shares 9,692 (-78.4%)

SEGALL BRYANT & HAMILL, LLC

Value ($000) $30,647 (-4.0%) Shares 921,984 (-7.8%)

DARK FOREST CAPITAL MANAGEMENT LP

Value ($000) $3,426 (-22.1%) Shares 103,059 (-25.2%)

EMERALD MUTUAL FUND ADVISERS TRUST

Value ($000) $3,021 (-14.4%) Shares 90,898 (-17.8%)

SJS Investment Consulting Inc.

Value ($000) $2,468 (-1.8%) Shares 74,236 (-5.7%)

SRS Capital Advisors, Inc.

Value ($000) $8 (-99.2%) Shares 236 (-99.2%)

EARNEST PARTNERS LLC

Value ($000) $58,107 (+1.0%) Shares 1,748,092 (-3.0%)

RENAISSANCE TECHNOLOGIES LLC

Value ($000) $7,329 (-45.8%) Shares 220,481 (-47.9%)

Founders Financial Alliance, LLC

Value ($000) $270 (-22.7%) Shares 8,128 (-25.8%)

ROYCE & ASSOCIATES LP

Value ($000) $1,495 (-39.9%) Shares 44,968 (-42.3%)

PEREGRINE CAPITAL MANAGEMENT LLC

Value ($000) $5,525 (-0.8%) Shares 166,214 (-4.7%)

Trexquant Investment LP

Value ($000) $1,258 (-36.5%) Shares 37,832 (-39.0%)

CITADEL ADVISORS LLC

Value ($000) $3,383 (-77.7%) Shares 101,784 (-78.5%)

Sanctuary Advisors, LLC

Value ($000) $207 (-80.8%) Shares 6,231 (-81.9%)

Clearstead Advisors, LLC

Value ($000) $268 (-64.1%) Shares 8,050 (-65.5%)

Institution	Prev Shares	Curr Shares	Change
FJ Capital Management LLC	654,296	347,136	-307,160 (-46.9%)
MALTESE CAPITAL MANAGEMENT LLC	401,981	275,000	-126,981 (-31.6%)
QSV Equity Investors LLC	51,514	34,246	-17,268 (-33.5%)
BRANT POINT INVESTMENT MANAGEMENT LLC	294,095	204,105	-89,990 (-30.6%)
Bridge City Capital, LLC	95,986	82,700	-13,286 (-13.8%)
BFSG, LLC	44,799	9,692	-35,107 (-78.4%)
SEGALL BRYANT & HAMILL, LLC	1,000,100	921,984	-78,116 (-7.8%)
DARK FOREST CAPITAL MANAGEMENT LP	137,761	103,059	-34,702 (-25.2%)
EMERALD MUTUAL FUND ADVISERS TRUST	110,529	90,898	-19,631 (-17.8%)
SJS Investment Consulting Inc.	78,703	74,236	-4,467 (-5.7%)
SRS Capital Advisors, Inc.	31,250	236	-31,014 (-99.2%)
EARNEST PARTNERS LLC	1,802,452	1,748,092	-54,360 (-3.0%)
RENAISSANCE TECHNOLOGIES LLC	423,147	220,481	-202,666 (-47.9%)
Founders Financial Alliance, LLC	10,947	8,128	-2,819 (-25.8%)
ROYCE & ASSOCIATES LP	77,931	44,968	-32,963 (-42.3%)
PEREGRINE CAPITAL MANAGEMENT LLC	174,396	166,214	-8,182 (-4.7%)
Trexquant Investment LP	62,049	37,832	-24,217 (-39.0%)
CITADEL ADVISORS LLC	474,138	101,784	-372,354 (-78.5%)
Sanctuary Advisors, LLC	34,421	6,231	-28,190 (-81.9%)
Clearstead Advisors, LLC	23,316	8,050	-15,266 (-65.5%)

Exited Positions (43)

Fourthstone LLC

Value ($000) $7,141 Shares 223,648

MENDON CAPITAL ADVISORS CORP

Value ($000) $1,597 Shares 50,000

MACQUARIE GROUP LTD

Value ($000) $115,519 Shares 3,617,873

Empowered Funds, LLC

Value ($000) $10,597 Shares 331,876

Avalon Advisory Group

Value ($000) $356 Shares 11,138

FOX RUN MANAGEMENT, L.L.C.

Value ($000) $774 Shares 24,231

nVerses Capital, LLC

Value ($000) $80 Shares 2,700

Curi Capital, LLC

Value ($000) $1,604 Shares 50,227

Campbell & CO Investment Adviser LLC

Value ($000) $511 Shares 15,991

Aquatic Capital Management LLC

Value ($000) $1,097 Shares 34,354

Entropy Technologies, LP

Value ($000) $440 Shares 13,790

Numerai GP LLC

Value ($000) $208 Shares 6,510

Hennion & Walsh Asset Management, Inc.

Value ($000) $618 Shares 19,350

Winthrop Partners - WNY, LLC

Value ($000) $9,342 Shares 250

Bridgewater Associates, LP

Value ($000) $3,450 Shares 108,045

Quantbot Technologies LP

Value ($000) $559 Shares 17,504

Waldron Private Wealth LLC

Value ($000) $421 Shares 13,021

Schonfeld Strategic Advisors LLC

Value ($000) $1,761 Shares 55,167

Elo Mutual Pension Insurance Co

Value ($000) $657 Shares 20,565

BAILARD, INC.

Value ($000) $283 Shares 8,859

Institution	Prev Shares	Prev Value ($000)
Fourthstone LLC	223,648	$7,141
MENDON CAPITAL ADVISORS CORP	50,000	$1,597
MACQUARIE GROUP LTD	3,617,873	$115,519
Empowered Funds, LLC	331,876	$10,597
Avalon Advisory Group	11,138	$356
FOX RUN MANAGEMENT, L.L.C.	24,231	$774
nVerses Capital, LLC	2,700	$80
Curi Capital, LLC	50,227	$1,604
Campbell & CO Investment Adviser LLC	15,991	$511
Aquatic Capital Management LLC	34,354	$1,097
Entropy Technologies, LP	13,790	$440
Numerai GP LLC	6,510	$208
Hennion & Walsh Asset Management, Inc.	19,350	$618
Winthrop Partners - WNY, LLC	250	$9,342
Bridgewater Associates, LP	108,045	$3,450
Quantbot Technologies LP	17,504	$559
Waldron Private Wealth LLC	13,021	$421
Schonfeld Strategic Advisors LLC	55,167	$1,761
Elo Mutual Pension Insurance Co	20,565	$657
BAILARD, INC.	8,859	$283