NAUTILUS BIOTECHNOLOGY INC

Ticker: NAUT CUSIP: 63909J108 Class: Common Stock

Note: Option positions (PUT/CALL) are excluded.

InstitutionSharesValue ($000)
a16z Capital Management, L.L.C.17,653,917$34,425
PERCEPTIVE ADVISORS LLC12,594,211$24,559
Cercano Management LLC7,172,985$13,987
Madrona Venture Group, LLC6,020,770$12
VANGUARD GROUP INC2,598,223$5,067
Comprehensive Financial Management LLC2,207,619$4,305
AMAZON COM INC1,457,055$2,841
Clarius Group, LLC1,209,646$2,359
MILLENNIUM MANAGEMENT LLC1,019,996$1,989
SENTINEL TRUST CO LBA750,000$1
Tikvah Management LLC726,895$1,417
ACADIAN ASSET MANAGEMENT LLC611,984$1
GEODE CAPITAL MANAGEMENT, LLC560,909$1,094
GOLDMAN SACHS GROUP INC297,437$580
DIMENSIONAL FUND ADVISORS LP285,439$556
RENAISSANCE TECHNOLOGIES LLC259,900$507
Alyeska Investment Group, L.P.253,468$494
JANE STREET GROUP, LLC206,837$403
STATE STREET CORP201,255$392
HILLMAN CO200,000$390
BlackRock, Inc.189,431$369
Crewe Advisors LLC136,052$265
BRIDGEWAY CAPITAL MANAGEMENT, LLC130,000$254
GSA CAPITAL PARTNERS LLP124,051$0
Qube Research & Technologies Ltd123,121$240
MARSHALL WACE, LLP117,059$228
Corient Private Wealth LLC114,788$224
NORTHERN TRUST CORP107,011$209
OMERS ADMINISTRATION Corp104,785$204
CITIGROUP INC61,712$120
Point72 Asset Management, L.P.60,833$119
HSBC HOLDINGS PLC49,449$98
BARCLAYS PLC48,254$94
Fayez Sarofim & Co45,000$88
Ashton Thomas Private Wealth, LLC45,000$88
Caprock Group, LLC44,090$86
SUSQUEHANNA INTERNATIONAL GROUP, LLP42,916$84
Palumbo Wealth Management LLC41,133$80
Lighthouse Investment Partners, LLC40,200$78
MORGAN STANLEY38,584$75
Presidio Capital Management, LLC34,664$68
CAPTRUST FINANCIAL ADVISORS34,011$66
XTX Topco Ltd33,608$66
Badgley Phelps Wealth Managers, LLC30,000$59
BNP PARIBAS ARBITRAGE, SA27,711$54
Callan Family Office, LLC25,970$51
Virtu Financial LLC23,532$0
BANK OF AMERICA CORP /DE/20,167$39
Hudson Bay Capital Management LP19,062$37
Garde Capital, Inc.18,000$35
TWO SIGMA SECURITIES, LLC16,239$32
FNY Investment Advisers, LLC14,000$0
Vanguard Capital Wealth Advisors13,000$25
Squarepoint Ops LLC11,316$22
UBS Group AG5,163$10
Rockefeller Capital Management L.P.5,000$10
GROUP ONE TRADING, L.P.3,880$8
FMR LLC3,486$7
ROYAL BANK OF CANADA3,274$6
TD Waterhouse Canada Inc.1,000$2
OSAIC HOLDINGS, INC.715$1
WELLS FARGO & COMPANY/MN560$1
Your Advocates Ltd., LLP150$0
CWM, LLC55$0
EverSource Wealth Advisors, LLC49$0
Optiver Holding B.V.40$0
Covestor Ltd8$0
Tower Research Capital LLC (TRC)5$0
Total (68 institutions)58,296,680$98,983