LIGAND PHARMACEUTICALS INC

Ticker: LGND CUSIP: 53220K504 Class: Common Stock

Note: Option positions (PUT/CALL) are excluded.

InstitutionSharesValue ($000)
BlackRock, Inc.2,940,231$555,909
VANGUARD GROUP INC2,073,892$392,111
JANUS HENDERSON GROUP PLC995,400$188,199
STATE STREET CORP751,205$142,030
NOMURA ASSET MANAGEMENT INTERNATIONAL INC.644,093$121,779
FMR LLC579,155$109,501
Chicago Capital, LLC561,423$106,148
Invesco Ltd.516,464$97,648
MASSACHUSETTS FINANCIAL SERVICES CO /MA/507,019$95,862
Stephens Investment Management Group LLC501,040$94,732
FRANKLIN RESOURCES INC496,160$93,809
GEODE CAPITAL MANAGEMENT, LLC493,589$93,336
CONGRESS ASSET MANAGEMENT CO /MA480,063$90,766
DIMENSIONAL FUND ADVISORS LP432,992$81,867
GOLDMAN SACHS GROUP INC429,004$81,112
LOOMIS SAYLES & CO L P382,677$72,353
UBS ASSET MANAGEMENT AMERICAS INC363,584$68,743
DEUTSCHE BANK AG\307,871$58,209
RICE HALL JAMES & ASSOCIATES, LLC279,627$52,869
Ranger Investment Management, L.P.269,983$51,046
VICTORY CAPITAL MANAGEMENT INC256,410$48,479
NORTHERN TRUST CORP250,640$47,389
ASHFORD CAPITAL MANAGEMENT INC213,778$40,419
Private Capital Management, LLC208,891$39,495
VILLERE ST DENIS J & CO LLC202,452$38,278
MORGAN STANLEY193,402$36,567
CHARLES SCHWAB INVESTMENT MANAGEMENT INC184,284$34,843
Impax Asset Management Group plc165,208$31,236
UBS Group AG143,067$27,050
Russell Investments Group, Ltd.138,464$26,179
WELLS FARGO & COMPANY/MN132,636$25,077
Fiera Capital Corp131,236$24,813
Fisher Asset Management, LLC130,858$24,741
JACOBS LEVY EQUITY MANAGEMENT, INC126,754$23,965
BANK OF MONTREAL /CAN/126,063$23,835
DRIEHAUS CAPITAL MANAGEMENT LLC122,326$23,128
Bank of New York Mellon Corp122,265$23,117
BANK OF AMERICA CORP /DE/120,169$22,720
abrdn plc119,309$22,558
GW&K Investment Management, LLC117,432$22
NOMURA HOLDINGS INC116,417$22,011
ROYAL BANK OF CANADA108,796$20,569
Legal & General Group Plc108,383$20,492
NEW YORK STATE COMMON RETIREMENT FUND107,530$20,331
Penn Capital Management Company, LLC98,848$18,710
AMERIPRISE FINANCIAL INC98,264$18,579
PRINCIPAL FINANCIAL GROUP INC93,830$17,740
Clark Capital Management Group, Inc.79,778$15,084
Allspring Global Investments Holdings, LLC71,818$13,653
Dana Investment Advisors, Inc.69,396$13,121
FIRST TRUST ADVISORS LP69,220$13,087
Public Sector Pension Investment Board68,114$12,878
CASTLEARK MANAGEMENT LLC66,736$12,618
Pictet Asset Management Holding SA64,386$12,547
KORNITZER CAPITAL MANAGEMENT INC /KS60,342$11,409
ESSEX INVESTMENT MANAGEMENT CO LLC60,020$11,348
OSAIC HOLDINGS, INC.59,405$11,232
Granite Investment Partners, LLC57,926$10,952
RHUMBLINE ADVISERS56,736$10,727
JPMORGAN CHASE & CO56,590$10,699
NORDEA INVESTMENT MANAGEMENT AB53,725$10,189
SEI INVESTMENTS CO46,368$8,767
F/M Investments LLC46,130$8,722
PRUDENTIAL FINANCIAL INC45,636$8,628
WCM INVESTMENT MANAGEMENT, LLC45,621$8,677
Lisanti Capital Growth, LLC43,742$8,270
Uniplan Investment Counsel, Inc.42,364$8,010
AMERICAN CENTURY COMPANIES INC40,885$7,730
Swiss National Bank38,600$7,298
Nuveen, LLC37,599$7,109
D. E. Shaw & Co., Inc.36,551$6,911
BNP PARIBAS ARBITRAGE, SA34,957$6,609
BARCLAYS PLC32,982$6,236
ENVESTNET ASSET MANAGEMENT INC27,735$5,244
NEW YORK STATE TEACHERS RETIREMENT SYSTEM26,950$5
Granahan Investment Management, LLC26,050$4,925
CIBC WORLD MARKET INC.25,630$4,846
ALLIANCEBERNSTEIN L.P.24,171$4,570
BESSEMER GROUP INC23,510$4
Edgestream Partners, L.P.22,889$4,328
EAM Investors, LLC21,092$3,988
Calamos Advisors LLC21,045$3,979
Universal- Beteiligungs- und Servicegesellschaft mbH20,902$3,952
ARROWSTREET CAPITAL, LIMITED PARTNERSHIP20,036$3,788
WOODSTOCK CORP20,005$3,782
PRICE T ROWE ASSOCIATES INC /MD/19,992$4
Moody Aldrich Partners LLC19,581$3,702
CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM18,797$3,554
RAYMOND JAMES FINANCIAL INC18,756$3,546
STIFEL FINANCIAL CORP18,722$3,540
CITIGROUP INC18,292$3,458
GREAT WEST LIFE ASSURANCE CO /CAN/16,750$3
Linden Thomas Advisory Services, LLC16,575$3,134
JANE STREET GROUP, LLC15,962$3,018
Qube Research & Technologies Ltd15,393$2,910
MARTINGALE ASSET MANAGEMENT L P15,322$2,897
CAPTRUST FINANCIAL ADVISORS14,579$2,756
CAXTON ASSOCIATES LLP14,508$2,743
State of Alaska, Department of Revenue13,649$3
Thrivent Financial for Lutherans13,525$3
244 other institutions635,708$107,986
Total (344 institutions)20,494,937$3,819,550