| Casdin Capital, LLC | 5,957,165 | $144,044 |
| BlackRock, Inc. | 5,911,614 | $142,943 |
| Invesco Ltd. | 3,932,191 | $95,080 |
| VANGUARD GROUP INC | 3,087,386 | $74,653 |
| FMR LLC | 1,548,194 | $37,435 |
| STATE STREET CORP | 1,505,593 | $36,405 |
| FRED ALGER MANAGEMENT, LLC | 1,083,838 | $26,207 |
| DIMENSIONAL FUND ADVISORS LP | 1,025,170 | $24,791 |
| MASSACHUSETTS FINANCIAL SERVICES CO /MA/ | 1,004,164 | $24,281 |
| GEODE CAPITAL MANAGEMENT, LLC | 953,126 | $23,050 |
| AMERIPRISE FINANCIAL INC | 904,318 | $21,866 |
| ROYAL BANK OF CANADA | 819,288 | $19,812 |
| PALISADE CAPITAL MANAGEMENT, LP | 811,010 | $19,610 |
| GENEVA CAPITAL MANAGEMENT LLC | 803,538 | $19,430 |
| Rockefeller Capital Management L.P. | 705,863 | $17,068 |
| Stephens Investment Management Group LLC | 606,887 | $14,675 |
| MUTUAL OF AMERICA CAPITAL MANAGEMENT LLC | 575,463 | $13,915 |
| MORGAN STANLEY | 535,040 | $12,937 |
| GAGNON SECURITIES LLC | 534,357 | $12,921 |
| BANK OF AMERICA CORP /DE/ | 521,327 | $12,606 |
| Voya Investment Management LLC | 496,937 | $12,016 |
| Granahan Investment Management, LLC | 482,356 | $11,663 |
| RAYMOND JAMES FINANCIAL INC | 440,054 | $10,826 |
| Curi Capital, LLC | 423,542 | $10,241 |
| NORTHERN TRUST CORP | 418,678 | $10,124 |
| ROYCE & ASSOCIATES LP | 417,090 | $10,085 |
| CHARLES SCHWAB INVESTMENT MANAGEMENT INC | 379,741 | $9,182 |
| Capricorn Fund Managers Ltd | 378,550 | $9,153 |
| PEREGRINE CAPITAL MANAGEMENT LLC | 356,343 | $8,616 |
| NEW YORK STATE COMMON RETIREMENT FUND | 298,618 | $7,221 |
| T. Rowe Price Investment Management, Inc. | 276,961 | $7 |
| Gagnon Advisors, LLC | 274,217 | $6,631 |
| Ranger Investment Management, L.P. | 270,290 | $6,536 |
| NEXT CENTURY GROWTH INVESTORS LLC | 262,971 | $6,359 |
| JPMORGAN CHASE & CO | 243,560 | $5,889 |
| UBS Group AG | 243,305 | $5,883 |
| GOLDMAN SACHS GROUP INC | 228,941 | $5,536 |
| RENAISSANCE TECHNOLOGIES LLC | 224,700 | $5,433 |
| ENVESTNET ASSET MANAGEMENT INC | 223,922 | $5,414 |
| Heights Point Management, LP | 217,548 | $5,260 |
| Bank of New York Mellon Corp | 216,283 | $5,230 |
| Uniplan Investment Counsel, Inc. | 194,617 | $4,706 |
| PRINCIPAL FINANCIAL GROUP INC | 186,613 | $4,512 |
| TD Asset Management Inc | 170,970 | $4,134 |
| D. E. Shaw & Co., Inc. | 166,381 | $4,023 |
| Russell Investments Group, Ltd. | 166,229 | $4,019 |
| ISTHMUS PARTNERS, LLC | 138,393 | $3,346 |
| Public Sector Pension Investment Board | 134,935 | $3,263 |
| WELLS FARGO & COMPANY/MN | 128,970 | $3,118 |
| SEI INVESTMENTS CO | 121,105 | $2,928 |
| DEERFIELD MANAGEMENT COMPANY, L.P. | 120,211 | $2,907 |
| Nuveen, LLC | 111,956 | $2,707 |
| Legal & General Group Plc | 105,014 | $2,539 |
| NOMURA ASSET MANAGEMENT INTERNATIONAL INC. | 99,579 | $2,408 |
| STATE OF WISCONSIN INVESTMENT BOARD | 98,213 | $2,375 |
| CITADEL ADVISORS LLC | 97,313 | $2,353 |
| RHUMBLINE ADVISERS | 93,296 | $2,256 |
| BARINGS LLC | 91,316 | $2,208 |
| Swiss National Bank | 80,300 | $1,942 |
| Conestoga Capital Advisors, LLC | 78,492 | $1,898 |
| Rathbones Group PLC | 77,086 | $1,864 |
| UBS ASSET MANAGEMENT AMERICAS INC | 75,372 | $1,822 |
| SummitTX Capital, L.P. | 72,266 | $1,747 |
| ZACKS INVESTMENT MANAGEMENT | 69,316 | $1,676 |
| BARCLAYS PLC | 64,094 | $1,550 |
| NEW YORK STATE TEACHERS RETIREMENT SYSTEM | 57,569 | $1 |
| FIRST TRUST ADVISORS LP | 50,354 | $1,218 |
| Cerity Partners LLC | 44,446 | $1,075 |
| KORNITZER CAPITAL MANAGEMENT INC /KS | 44,188 | $1,068 |
| ALLIANCEBERNSTEIN L.P. | 44,012 | $1,064 |
| JANE STREET GROUP, LLC | 43,966 | $1,063 |
| CITIGROUP INC | 40,979 | $991 |
| CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM | 38,733 | $937 |
| CIBC Private Wealth Group, LLC | 37,697 | $912 |
| D.A. DAVIDSON & CO. | 37,556 | $908 |
| XTX Topco Ltd | 36,262 | $877 |
| PRUDENTIAL FINANCIAL INC | 35,752 | $864 |
| GREAT WEST LIFE ASSURANCE CO /CAN/ | 34,692 | $1 |
| Qube Research & Technologies Ltd | 33,410 | $808 |
| SANDLER CAPITAL MANAGEMENT | 31,637 | $765 |
| GREAT LAKES ADVISORS, LLC | 30,919 | $748 |
| PRICE T ROWE ASSOCIATES INC /MD/ | 29,783 | $1 |
| DEUTSCHE BANK AG\ | 29,694 | $718 |
| Meros Investment Management, LP | 28,454 | $688 |
| SG Americas Securities, LLC | 27,128 | $1 |
| Thrivent Financial for Lutherans | 26,798 | $1 |
| Diametric Capital, LP | 25,622 | $620 |
| INTECH INVESTMENT MANAGEMENT LLC | 25,571 | $618 |
| Cannell & Spears LLC | 25,436 | $615 |
| Allspring Global Investments Holdings, LLC | 24,622 | $595 |
| Penbrook Management LLC | 22,955 | $555 |
| Illinois Municipal Retirement Fund | 22,662 | $548 |
| MetLife Investment Management, LLC | 22,042 | $533 |
| Campbell & CO Investment Adviser LLC | 21,820 | $528 |
| State of Alaska, Department of Revenue | 19,850 | $0 |
| JANUS HENDERSON GROUP PLC | 19,717 | $476 |
| MANUFACTURERS LIFE INSURANCE COMPANY, THE | 19,221 | $465 |
| MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd. | 19,062 | $461 |
| BNP PARIBAS ARBITRAGE, SA | 18,960 | $458 |
| Louisiana State Employees Retirement System | 17,900 | $433 |
| 85 other institutions | 355,199 | $7,746 |