| VANGUARD GROUP INC | 9,101,779 | $45,509 |
| BlackRock, Inc. | 9,026,700 | $45,134 |
| Granahan Investment Management, LLC | 6,349,463 | $31,747 |
| BANK OF AMERICA CORP /DE/ | 3,084,002 | $15,420 |
| STATE STREET CORP | 2,789,126 | $13,946 |
| GEODE CAPITAL MANAGEMENT, LLC | 2,528,772 | $12,646 |
| ARROWSTREET CAPITAL, LIMITED PARTNERSHIP | 2,129,363 | $10,647 |
| Manatuck Hill Partners, LLC | 2,000,000 | $10,000 |
| Invesco Ltd. | 1,829,003 | $9,145 |
| DIMENSIONAL FUND ADVISORS LP | 1,507,478 | $7,538 |
| MORGAN STANLEY | 1,427,510 | $7,138 |
| G2 Investment Partners Management LLC | 1,404,079 | $7,020 |
| TWO SIGMA INVESTMENTS, LP | 1,246,343 | $6,232 |
| D. E. Shaw & Co., Inc. | 1,163,178 | $5,816 |
| CITADEL ADVISORS LLC | 1,074,205 | $5,371 |
| SILVERCREST ASSET MANAGEMENT GROUP LLC | 1,041,060 | $5,205 |
| JANE STREET GROUP, LLC | 959,412 | $4,797 |
| AQR CAPITAL MANAGEMENT LLC | 928,392 | $4,642 |
| Squarepoint Ops LLC | 919,027 | $4,595 |
| BARCLAYS PLC | 859,408 | $4,297 |
| CHARLES SCHWAB INVESTMENT MANAGEMENT INC | 852,731 | $4,264 |
| NORTHERN TRUST CORP | 820,336 | $4,102 |
| HRT FINANCIAL LP | 810,647 | $4 |
| UBS Group AG | 779,585 | $3,898 |
| RENAISSANCE TECHNOLOGIES LLC | 742,300 | $3,712 |
| HARBOR CAPITAL ADVISORS, INC. | 692,459 | $3 |
| GOLDMAN SACHS GROUP INC | 663,773 | $3,319 |
| S Squared Technology, LLC | 653,571 | $3,268 |
| Counterpoint Mutual Funds LLC | 634,542 | $3,173 |
| Herald Investment Management Ltd | 600,000 | $2,994 |
| ACADIAN ASSET MANAGEMENT LLC | 596,145 | $3 |
| JPMORGAN CHASE & CO | 567,520 | $2,838 |
| Trexquant Investment LP | 407,097 | $2,035 |
| GC Wealth Management RIA, LLC | 406,139 | $2,031 |
| Bank of New York Mellon Corp | 390,698 | $1,953 |
| TUDOR INVESTMENT CORP ET AL | 368,310 | $1,842 |
| PERKINS CAPITAL MANAGEMENT INC | 366,100 | $1,831 |
| Nuveen, LLC | 355,731 | $1,779 |
| BOOTHBAY FUND MANAGEMENT, LLC | 331,699 | $1,658 |
| CITIGROUP INC | 301,060 | $1,505 |
| RAYMOND JAMES FINANCIAL INC | 300,483 | $1,502 |
| P.A.W. CAPITAL CORP | 290,000 | $1,450 |
| Verition Fund Management LLC | 286,616 | $1,433 |
| Point72 Asset Management, L.P. | 267,519 | $1,338 |
| PRUDENTIAL FINANCIAL INC | 263,474 | $1,317 |
| Universal- Beteiligungs- und Servicegesellschaft mbH | 255,499 | $1,288 |
| Baader Bank Aktiengesellschaft | 252,436 | $1,262 |
| Soviero Asset Management, LP | 240,000 | $1,200 |
| Engineers Gate Manager LP | 209,720 | $1,049 |
| ALGERT GLOBAL LLC | 207,040 | $1 |
| XTX Topco Ltd | 196,075 | $980 |
| GSA CAPITAL PARTNERS LLP | 189,394 | $1 |
| Russell Investments Group, Ltd. | 186,924 | $935 |
| TWO SIGMA ADVISERS, LP | 184,374 | $922 |
| SG Americas Securities, LLC | 176,354 | $1 |
| WELLS FARGO & COMPANY/MN | 162,359 | $812 |
| HighTower Advisors, LLC | 145,413 | $727 |
| ALLIANCEBERNSTEIN L.P. | 140,970 | $705 |
| PRICE T ROWE ASSOCIATES INC /MD/ | 140,271 | $1 |
| ExodusPoint Capital Management, LP | 136,321 | $682 |
| RHUMBLINE ADVISERS | 132,994 | $665 |
| Jump Financial, LLC | 130,756 | $654 |
| MILLENNIUM MANAGEMENT LLC | 128,540 | $643 |
| FULLER & THALER ASSET MANAGEMENT, INC. | 128,420 | $642 |
| PANAGORA ASSET MANAGEMENT INC | 103,448 | $517 |
| DEUTSCHE BANK AG\ | 101,736 | $509 |
| Rathbones Group PLC | 101,015 | $505 |
| MACKENZIE FINANCIAL CORP | 99,772 | $502 |
| Quantbot Technologies LP | 95,179 | $476 |
| Round Rock Advisors, LLC | 93,689 | $468 |
| Worth Venture Partners, LLC | 85,125 | $426 |
| PharVision Advisers, LLC | 82,744 | $414 |
| GAGNON SECURITIES LLC | 81,839 | $409 |
| IEQ CAPITAL, LLC | 81,604 | $408 |
| SIMPLEX TRADING, LLC | 81,105 | $406 |
| NATIXIS | 79,300 | $397 |
| BNP PARIBAS ARBITRAGE, SA | 73,265 | $366 |
| CastleKnight Management LP | 70,100 | $351 |
| Brendel Financial Advisors LLC | 68,007 | $340 |
| Kingdom Financial Group LLC. | 67,066 | $335 |
| LPL Financial LLC | 62,458 | $312 |
| State of Alaska, Department of Revenue | 60,550 | $0 |
| BANK OF MONTREAL /CAN/ | 58,487 | $292 |
| HSBC HOLDINGS PLC | 56,994 | $288 |
| Mariner, LLC | 54,562 | $273 |
| MetLife Investment Management, LLC | 51,209 | $256 |
| FRANKLIN RESOURCES INC | 50,331 | $252 |
| MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd. | 50,200 | $251 |
| Stonehage Fleming Financial Services Holdings Ltd | 49,920 | $250 |
| FMR LLC | 47,960 | $240 |
| Sanctuary Advisors, LLC | 46,654 | $233 |
| ROYAL BANK OF CANADA | 46,526 | $232 |
| VARCOV Co. | 46,465 | $232 |
| BAILARD, INC. | 45,829 | $229 |
| CWM, LLC | 42,083 | $0 |
| STATE OF WISCONSIN INVESTMENT BOARD | 40,528 | $203 |
| IMC-Chicago, LLC | 40,047 | $200 |
| Lido Advisors, LLC | 39,256 | $251 |
| COMMONWEALTH EQUITY SERVICES, LLC | 37,627 | $188 |
| INTECH INVESTMENT MANAGEMENT LLC | 36,238 | $181 |
| 95 other institutions | 965,919 | $4,600 |