TRUPANION INC

Ticker: TRUP CUSIP: 898202106 Class: Common Stock

Note: Option positions (PUT/CALL) are excluded.

InstitutionSharesValue ($000)
BlackRock, Inc.5,382,374$201,139
Capital World Investors4,123,266$154,086
Aflac Inc3,636,364$135,891
VANGUARD GROUP INC3,505,021$130,983
Polar Capital Holdings Plc1,475,000$55,121
FMR LLC1,400,657$52,343
STATE STREET CORP1,290,987$48,244
MARSHALL WACE, LLP1,267,262$47,358
GEODE CAPITAL MANAGEMENT, LLC865,841$32,362
MORGAN STANLEY787,460$29,427
GREENLEA LANE CAPITAL MANAGEMENT, LLC776,490$29,017
DIMENSIONAL FUND ADVISORS LP607,022$22,686
BANK OF AMERICA CORP /DE/597,802$22,340
GOLDMAN SACHS GROUP INC477,014$17,826
Hosking Partners LLP465,950$17,413
Clearbridge Investments, LLC447,854$16,736
NORTHERN TRUST CORP368,631$13,776
CITADEL ADVISORS LLC332,800$12,437
CHARLES SCHWAB INVESTMENT MANAGEMENT INC330,521$12,352
Trexquant Investment LP316,463$11,826
BARCLAYS PLC302,625$11,309
Kopion Asset Management, LLC293,706$10,976
MILLENNIUM MANAGEMENT LLC283,746$10,604
BALYASNY ASSET MANAGEMENT L.P.276,580$10,336
PRINCIPAL FINANCIAL GROUP INC268,203$10,023
RENAISSANCERE HOLDINGS LTD263,000$9,828
Lagoda Investment Management, L.P.249,275$9,315
WELLINGTON MANAGEMENT GROUP LLP248,780$9,297
D. E. Shaw & Co., Inc.247,312$9,242
TWO SIGMA INVESTMENTS, LP205,055$7,663
Bank of New York Mellon Corp196,068$7,327
Qube Research & Technologies Ltd174,231$6,511
Public Sector Pension Investment Board170,121$6,357
BNP PARIBAS ARBITRAGE, SA168,037$6,280
Allianz Asset Management GmbH164,884$6,162
TUDOR INVESTMENT CORP ET AL138,687$5,183
MAVERICK CAPITAL LTD133,873$5,003
CITIGROUP INC131,244$4,905
Engineers Gate Manager LP130,791$4,888
HEALTHCARE OF ONTARIO PENSION PLAN TRUST FUND124,329$4,646
WELLS FARGO & COMPANY/MN122,935$4,594
PDT Partners, LLC119,577$4,469
Invesco Ltd.112,076$4,188
UBS Group AG106,049$3,963
Squarepoint Ops LLC102,255$3,821
Nuveen, LLC101,625$3,798
RHUMBLINE ADVISERS95,770$3,579
Legal & General Group Plc87,642$3,275
HRT FINANCIAL LP83,123$3
TWO SIGMA ADVISERS, LP75,300$2,814
FIRST TRUST ADVISORS LP73,633$2,752
CONNORS INVESTOR SERVICES INC71,744$2,681
AMERICAN CENTURY COMPANIES INC71,506$2,672
DEUTSCHE BANK AG\69,936$2,614
DEERFIELD MANAGEMENT COMPANY, L.P.66,438$2,483
UBS ASSET MANAGEMENT AMERICAS INC66,368$2,480
SUSQUEHANNA FUNDAMENTAL INVESTMENTS, LLC66,368$2,480
AMERIPRISE FINANCIAL INC64,556$2,412
ProShare Advisors LLC63,895$2,388
Swiss National Bank63,610$2,377
INGALLS & SNYDER LLC62,500$2
ALLIANCEBERNSTEIN L.P.51,104$1,910
Liontrust Investment Partners LLP48,992$1,831
Universal- Beteiligungs- und Servicegesellschaft mbH48,987$1,868
CHECK CAPITAL MANAGEMENT INC/CA40,780$1,524
Dynamic Technology Lab Private Ltd37,329$1,395
JANE STREET GROUP, LLC37,287$1,393
NEW YORK STATE TEACHERS RETIREMENT SYSTEM34,567$1
Verition Fund Management LLC34,200$1,278
PRICE T ROWE ASSOCIATES INC /MD/33,657$1
PRUDENTIAL FINANCIAL INC32,005$1,196
Blueshift Asset Management, LLC31,997$1,196
CALIFORNIA STATE TEACHERS RETIREMENT SYSTEM31,358$1,172
GREAT WEST LIFE ASSURANCE CO /CAN/29,751$1
Atom Investors LP29,617$1,107
JPMORGAN CHASE & CO29,585$1,106
JACOBS LEVY EQUITY MANAGEMENT, INC29,510$1,103
WINTON GROUP Ltd28,271$1,056
CANADA PENSION PLAN INVESTMENT BOARD25,900$968
Saranac Partners Ltd24,420$913
Thrivent Financial for Lutherans23,153$1
LOS ANGELES CAPITAL MANAGEMENT LLC23,000$931
FOX RUN MANAGEMENT, L.L.C.21,828$816
New Age Alpha Advisors, LLC21,367$798
MetLife Investment Management, LLC20,884$780
State of Wyoming20,189$754
AQR CAPITAL MANAGEMENT LLC20,159$753
ROYAL BANK OF CANADA19,769$738
INTECH INVESTMENT MANAGEMENT LLC17,726$662
Russell Investments Group, Ltd.17,499$654
Illinois Municipal Retirement Fund17,413$651
MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd.17,259$645
State of Alaska, Department of Revenue17,156$1
Jump Financial, LLC16,700$624
MANUFACTURERS LIFE INSURANCE COMPANY, THE16,209$606
Louisiana State Employees Retirement System15,400$575
JANUS HENDERSON GROUP PLC14,940$558
AdvisorNet Financial, Inc14,760$552
RAYMOND JAMES FINANCIAL INC14,617$546
Woodline Partners LP14,498$542
112 other institutions446,189$14,799
Total (212 institutions)35,842,264$1,327,066