| NEEDHAM INVESTMENT MANAGEMENT LLC | 3,411,500 | $52,878 |
| BlackRock, Inc. | 2,016,987 | $31,263 |
| VANGUARD GROUP INC | 1,852,669 | $28,716 |
| FRANKLIN RESOURCES INC | 1,743,031 | $27,017 |
| WESTERLY CAPITAL MANAGEMENT, LLC | 1,295,000 | $20,073 |
| Samjo Management, LLC | 1,027,130 | $15,921 |
| GOLDMAN SACHS GROUP INC | 1,012,670 | $15,696 |
| ACADIAN ASSET MANAGEMENT LLC | 912,290 | $14 |
| GEODE CAPITAL MANAGEMENT, LLC | 760,387 | $11,789 |
| STATE STREET CORP | 760,113 | $11,782 |
| Nuveen, LLC | 737,281 | $11,428 |
| SILVERCREST ASSET MANAGEMENT GROUP LLC | 512,922 | $7,950 |
| NEXT CENTURY GROWTH INVESTORS LLC | 493,367 | $7,647 |
| AQR CAPITAL MANAGEMENT LLC | 490,044 | $7,596 |
| FEDERATED HERMES, INC. | 437,608 | $6,783 |
| ALGERT GLOBAL LLC | 402,912 | $6 |
| G2 Investment Partners Management LLC | 400,109 | $6,202 |
| Herald Investment Management Ltd | 400,000 | $6,192 |
| JANE STREET GROUP, LLC | 380,818 | $5,903 |
| Trexquant Investment LP | 378,893 | $5,873 |
| MILLENNIUM MANAGEMENT LLC | 375,094 | $5,814 |
| EAM Investors, LLC | 371,276 | $5,755 |
| THAMES CAPITAL MANAGEMENT LLC | 293,973 | $4,557 |
| STATE OF WISCONSIN INVESTMENT BOARD | 288,545 | $4,472 |
| ESSEX INVESTMENT MANAGEMENT CO LLC | 285,167 | $4,420 |
| MORGAN STANLEY | 244,695 | $3,793 |
| Granahan Investment Management, LLC | 236,896 | $3,672 |
| NORTHERN TRUST CORP | 227,836 | $3,531 |
| JACOBS LEVY EQUITY MANAGEMENT, INC | 224,695 | $3,483 |
| MARSHALL WACE, LLP | 209,782 | $3,252 |
| LPL Financial LLC | 163,464 | $2,534 |
| TUDOR INVESTMENT CORP ET AL | 159,576 | $2,473 |
| Squarepoint Ops LLC | 153,492 | $2,379 |
| Point72 Asset Management, L.P. | 143,704 | $2,227 |
| NANO CAP NEW MILLENNIUM GROWTH FUND L P | 123,500 | $1,914 |
| SEI INVESTMENTS CO | 118,993 | $1,844 |
| Huntleigh Advisors, Inc. | 118,220 | $1,832 |
| PANAGORA ASSET MANAGEMENT INC | 112,348 | $1,741 |
| BOOTHBAY FUND MANAGEMENT, LLC | 109,134 | $1,692 |
| Bridgewater Associates, LP | 104,081 | $1,613 |
| JANUS HENDERSON GROUP PLC | 96,859 | $1,501 |
| CHARLES SCHWAB INVESTMENT MANAGEMENT INC | 94,888 | $1,471 |
| RENAISSANCE TECHNOLOGIES LLC | 88,700 | $1,375 |
| Bank of New York Mellon Corp | 74,720 | $1,158 |
| IEQ CAPITAL, LLC | 71,210 | $1,104 |
| UBS Group AG | 70,216 | $1,088 |
| AMERIPRISE FINANCIAL INC | 64,757 | $1,004 |
| CITADEL ADVISORS LLC | 63,143 | $979 |
| BANK OF AMERICA CORP /DE/ | 55,643 | $862 |
| Mariner, LLC | 53,629 | $832 |
| WHITE PINE CAPITAL LLC | 52,991 | $821 |
| Russell Investments Group, Ltd. | 50,395 | $781 |
| RHUMBLINE ADVISERS | 47,663 | $739 |
| BARCLAYS PLC | 47,299 | $733 |
| SUSQUEHANNA INTERNATIONAL GROUP, LLP | 46,225 | $716 |
| FMR LLC | 41,912 | $650 |
| SIMPLEX TRADING, LLC | 39,718 | $616 |
| WELLS FARGO & COMPANY/MN | 36,662 | $568 |
| AlphaQuest LLC | 35,273 | $547 |
| Susquehanna Portfolio Strategies, LLC | 32,958 | $511 |
| GSA CAPITAL PARTNERS LLP | 32,500 | $1 |
| JPMORGAN CHASE & CO | 30,055 | $466 |
| Connor, Clark & Lunn Investment Management Ltd. | 29,854 | $463 |
| ARROWSTREET CAPITAL, LIMITED PARTNERSHIP | 29,707 | $460 |
| Qube Research & Technologies Ltd | 28,967 | $449 |
| XTX Topco Ltd | 27,334 | $424 |
| Graham Capital Management, L.P. | 26,488 | $411 |
| Numerai GP LLC | 25,129 | $390 |
| CWM, LLC | 24,129 | $0 |
| PUBLIC EMPLOYEES RETIREMENT SYSTEM OF OHIO | 23,913 | $371 |
| SG Americas Securities, LLC | 22,165 | $0 |
| DEUTSCHE BANK AG\ | 21,159 | $328 |
| PharVision Advisers, LLC | 20,655 | $320 |
| STRS OHIO | 20,500 | $318 |
| TWO SIGMA ADVISERS, LP | 18,400 | $285 |
| Dynamic Technology Lab Private Ltd | 17,586 | $273 |
| Invesco Ltd. | 17,499 | $271 |
| PRICE T ROWE ASSOCIATES INC /MD/ | 17,453 | $0 |
| MetLife Investment Management, LLC | 16,951 | $263 |
| WEALTH ENHANCEMENT ADVISORY SERVICES, LLC | 16,467 | $289 |
| Campbell & CO Investment Adviser LLC | 16,449 | $255 |
| Cerity Partners LLC | 15,055 | $233 |
| Quantinno Capital Management LP | 14,867 | $230 |
| D. E. Shaw & Co., Inc. | 14,649 | $227 |
| MIRAE ASSET GLOBAL ETFS HOLDINGS Ltd. | 14,106 | $219 |
| VestGen Advisors, LLC | 14,091 | $218 |
| INTECH INVESTMENT MANAGEMENT LLC | 13,540 | $210 |
| Creative Planning | 13,381 | $207 |
| BANK OF NOVA SCOTIA | 12,500 | $194 |
| State of Wyoming | 11,626 | $180 |
| SUSQUEHANNA FUNDAMENTAL INVESTMENTS, LLC | 11,200 | $174 |
| LOS ANGELES CAPITAL MANAGEMENT LLC | 11,050 | $147 |
| MERCER GLOBAL ADVISORS INC /ADV | 10,428 | $162 |
| NEW YORK STATE COMMON RETIREMENT FUND | 9,500 | $147 |
| Police & Firemen's Retirement System of New Jersey | 8,168 | $127 |
| BNP PARIBAS ARBITRAGE, SA | 7,294 | $113 |
| OSAIC HOLDINGS, INC. | 4,079 | $63 |
| CANADA PENSION PLAN INVESTMENT BOARD | 3,600 | $56 |
| Ameritas Investment Partners, Inc. | 2,703 | $42 |
| Legal & General Group Plc | 2,661 | $41 |
| 37 other institutions | 23,030 | $342 |